DANIEL & TRIPP: LOUTERIDIUM : TAXONOMY, BIOLOGY, AND CONSERVATION 91 
Daniel (1995) noted considerable variation in pubescence, leaf shape, bract persistence, and 
bract shape among specimens from Chiapas, Mexico. One of the two collections {Breedlove 24803, 
Palacios E. 385 ) that he indicated showed some intermediacy with L. donnell-smithii in pubes¬ 
cence was sampled in the phylogenetic analysis. Palacios E. 385 was strongly supported as nested 
within the clade of other sampled plants of L. mexicanum (Fig. 1). Thus, both plants are here 
included within L. mexicanum. 
Somewhat atypical plants from the vicinity of San Felipe Usila, Oaxaca (e.g., Ibarra M. et al. 
3758, 3765; Calzada et al. 16579 ) are distinctive by their long calyces (30-39 mm) with the pos¬ 
terior lobe conspicuously attenuate-falcate apically, corollas pale rose to white with purplish lobes 
and pubescent with glandular and eglandular trichomes to 0.2 mm long, and rachis (i.e., Calzada 
et al. 16579 ) often somewhat scurfy to minutely puberulent with sessile to subsessile glandular tri¬ 
chomes less than 0.05 mm long. In Calzada et al. 16579, the calyx and pedicels are like more 
typical specimens of L. mexicanum, and the rachis often has longer eglandular trichomes to 0.1 mm 
as well. Although similar calyces and trichomes on corollas are somet im es seen elsewhere among 
specimens of L. mexicanum, the corolla color of these collections appears to be unusual compared 
to other Mexican representatives of the species. However, corollas of Foerther et al. 10940 from 
Guatemala are described as brownish violet. In pubescence of the rachis, these plants somewhat 
resemble L. purpusii. On the whole, they appear more like L. mexicanum, in which species they are 
treated here. 
Additional specimens examined.— MEXICO. Chiapas: Mpio. Ocosingo, 4.4 km SO de 
San Javier, 16°48’49”N, 091°04’04 ,, W, G. Aguilar M. et al. 4713 (MEXU); Mpio. Solosuchiapa, 
2^1 km below Ixhuatan toward Pichucalco, [ca. 17°18’47.23”N, 093°00’11.57”W], D. Breedlove 
24177 (DS, ENCB, MO, NY), 29799 (DS, LL, MEXU, MICH, MO, RSA), D. Breedlove & 
F. Almeda 56805 (CAS, ENCB, LL, NY); Mpio. Berriozabal, 13 km N of Berriozabal near Pozo 
Turripache and Finca El Suspire, [ca. 16°52’25.01”N, 093°19’8.64”W], D. Breedlove 24803 (DS, 
ENCB, MO), D. Breedlove & T. Daniel 70879 (CAS), cultivated plants of this collection grown 
from cuttings in San Francisco, California, 70879cv (CAS), D. Breedlove & R. Thorne 30786 (DS, 
ENCB, MO), D. Breedlove et al. 66158 (CAS); Mpio. Palenque, near Agua Azul, D. Breedlove 
56091 (CAS, LL, MEXU); Mpio. Ocosingo, near ruins of Bonampak at jet. road from Lacanja to 
Echeverria and road from Chancala, [ca. 16°48’04.57”N, 091°06’21.97”W], D. Breedlove & 
F. Almeda 58020 (CAS, LL, MEXU, MO); Mpio. San Fernando, 20 km NW of San Fernando just 
beyond Colonia Cuahtemoc, [ca. 16°58’42.74”N, 093°20’28.46”W], D. Breedlove & M. Bourell 
68269 (CAS); Mpio. Ocosingo, 70 km SW of Palenque toward Ocosingo along the Jol Uk’um, [ca. 
17°10’48.02”N, 092°06’39.86”W], D. Breedlove & B. Keller 49526 (CAS, MEXU, MO); Mpio. 
Ocozocoautla de Espinosa, 26-28 km N of Ocozocoautla toward Mai Paso, [ca. 16°56’22.87”N, 
093°27’55.46”W], D. Breedlove & A. Smith 22469 (DS, MEXU, MICH, MO, RSA); Mpio. 
Ocosingo, 4 km E del crucero San Javier, 16°48’44”N, 091°03’13”W, J. Calonico S. et al. 25022 
(ENCB, MEXU); Mpio. Berriozabal, ca. 12 km NW of Berriozabal, ca. 16°51’N, 093°19’W, 
T. Daniel 8375 (CAS, MEXU); Mpio. Ishuatan, carretera 195, 45 km N [S] de Pichucalco, [ca. 
HHS^b.lT’N, 093°00’27.05”W], S. Gliessman 77-7 (MEXU); Mpio. Pichucalco, 20 km Teapa- 
Pichucalco, [ca. 17°32’45.83”N, 093°04’44.80”W], M. Ishiki et al. 2035 (CAS); Mpio. Ocosingo, 
Lacandona de Lacanha-Chansayab, 130 km SE de Palenque, por la carretera fronteriza hasta el 
crucero San Javier, despues 8 km hacia el oeste, 16°44’N, 091°05’W, S. Levy T. & A. Duran F. 398 
(MEXU); Mpio. Berriozabal, 12 km N de Berriozabal, E. Martinez S. & M. Soto 24216 (MEXU); 
Mpio. Ishuatan, 9 km N de Tapilula, [ca. 17 0 17’16.71”N, 093°00’38.70”W], E. Martinez S. et al. 
3198 (CAS, ENCB, MEXU, MO, WIS); Mpio. Yajalon, Yajalon-Petalsinco, carretera de terraceria 
KM 5, A. Mendez T. [=A. Ton] 7134 (IEB, MEXU, MO, RSA, XAL); Mpio. Ocozocoautla, 9 km 
