DANIEL & TRIPP: LOUTERIDIUM : TAXONOMY, BIOLOGY, AND CONSERVATION 97 
MEXU, MO, XAL); Mpio. Minatitlan, 6.6 km N de la terraceria La Langua-Rio Grande, sobre el 
camino nuevo a Ejido Belisario Dominguez, el cual sale de la terraceria 14.7 km E de La Laguna, 
17°20’N, 094°22’W, T. Wendt & A. Villalobos C. 2540 (CAS, ENCB, LSU, MEXU). 
9. Louteridium purpusii Brandegee, Univ. Calif. Publ. Bot. 6:68. 1914. Type.— MEXICO. 
Chiapas: Finca Irlanda [ca. 15°10’24.94”N, 092°20’ 10.22”W], VIII-1913 (fir, frt), C. Purpus 6969 
(holotype: UC!; isotypes: BM!, EAP-image seen, F!, GH!, MO!, NY!, US!, Z-image seen). 
Shrubs or trees to 9 m tall, terrestrial. Older (woody) stems subquadrate-sulcate, ± scurfy and 
lenticellate, often puberulent like young stems; younger (herbaceous) stems quadrate to quadrate- 
sulcate and ± irregularly striate, glabrous or ± evenly puberulent with erect to retrorse to antrorse 
eglandular trichomes to 0.05 mm long. Leaves apparently not all seasonally deciduous, ± evenly 
distributed along young stems, petiolate, petioles to 40 mm long, blades membranaceous, ovate- 
elliptic to obovate-elliptic to obovate, 130-380 mm long, 40-115 mm wide, 2.1—4.3 x longer than 
wide, acuminate to acute-apiculate at apex, acute to attenuate at base, adaxial surface glabrous or 
puberulent like young stems, abaxial surface paler than abaxial surface, glabrous or pubescent with 
mostly antrorse eglandular trichomes 0.05-0.1 mm long along major veins, margin often undulate, 
entire to sinuate to crenate to crenate-dentate. Inflorescence a terminal pedunculate raceme to race¬ 
mose thyrse to 63 cm long (including peduncles and excluding corollas), peduncle to 20 cm long, 
glabrous or pubescent like young stems, rachis puberulent with (retrorse to) erect to flexuose to 
antrorse eglandular trichomes to 0.05 mm long; dichasia modified by very short expansion between 
pairs of succeeding flowers with the congested dichasial axis becoming a ± fan-shaped to multi- 
and tortuously-branched racemelike lateral short-shoot to 15 mm long, opposite (and alternate?), 
sessile to subsessile, 3-many-flowered, to 60 mm long (excluding corollas), dichasial peduncles (if 
present) to 2 mm long, pubescent like rachis. Bracts caducous, not seen. Bracteoles and secondary 
bracteoles caducous, lance-linear to linear-elliptic, 5-13 mm long, 1-2.5 mm wide, abaxially 
pubescent like rachis. Flowers pedicellate, pedicels to 55 mm long, puberulent with erect to retrorse 
(to antrorse) eglandular trichomes to 0.05 mm long. Calyx 22^10 mm long, lobes distinct or fused 
at base up to 1 mm, heteromorphic, membranaceous, abaxially puberulent like pedicels and some¬ 
times with a few sparse stipitate glands up to 0.2 mm long as well, also inconspicuously glandu¬ 
lar-punctate, posterior lobe conduplicate, ovate to elliptic to obovate, 21-38 mm long, slightly 
shorter than to slightly longer than lateral lobes, 8.5-15 mm wide, acuminate to falcate at apex, 
lateral lobes sublunate, 22-37 mm long, 6-9 mm wide, acute to subfalcate at apex. Corolla cream 
to yellow-greenish and tinged with maroon, 45-52 mm long, externally pubescent with stipitate 
glandular trichomes 0.05-0.3 mm long (sometimes with flexuose eglandular trichomes to 0.2 mm 
long on or near margins of lobes as well), tube 25-33 mm long, narrow proximal portion 3-5 mm 
long, 7 mm in diameter near midpoint, throat 22-30 mm long, 25-35 mm diameter at mouth, lobes 
spreading to recurved, triangular to ovate, 12-21 mm long, 10-13 mm wide, rounded at apex. 
Stamens 2, 65-75 mm long, filaments glabrous distally, pubescent proximally with eglandular tri¬ 
chomes, thecae 10-12 mm long; staminodes (if present) not seen. Style 85-92 mm long, glabrous 
distally (not seen proximally), stigma equally 2-lobed, lobes elliptic, 1.4-2 mm long, width not 
determined. Capsule 22-30 mm long, 5-7.5 mm in diameter, densely pubescent with stipitate glan¬ 
dular trichomes (0.05-) 0.1-0.4 mm long, stipe 3-3.5 mm long. Seeds up to 12 per capsule, 
4.5-5.2 mm long, 4.5-5 mm wide, surfaces smooth. 
Phenology. — Flowering and fruiting: August-March. 
Distribution and habitat.— Southern Mexico (Chiapas), Guatemala (Fig. 10); plants occur 
in wooded ravines in montane rain forests on the Pacific versant at elevations from 600 to 2000 m. 
