GODDARD ET AL.: EL NINO IMPACT ON HETEROBRANCH SEA SLUG RANGES 113 
Figure 1. Heterobranch sea slugs found at new northernmost localities in the Northeastern Pacific Ocean, 2015-2017. 
A Okenia angelensis, Miwok Beach, Sonoma Co., California, 27 May 2017. Image by Colby Davidson. B Acanthodoris 
rhodoceras, Chup Point, Barkley Sound, Vancouver Island, British Columbia, 21 May 2018. Image by Peter Mieras/subvi- 
sionproductions.com. C Polycera atra, Lemmens Inlet, Clayoquot Sound, Vancouver Island, British Columbia, 25 July 
2015. Inset: tail. Images by Brandon Exner. D Thordisa rubescens, Santa Cruz Island, California, 1 November 2017. Image 
by Kenan Chan/Channel Islands National Park. 
laying eggs, the past two years on subtidal reefs around the Palos Verdes Peninsula (P. Garner, per¬ 
sonal communication to RA, 20 June 2018; and see <https://www.inaturalist.org/taxa/50139 
-Thordisa-rubescens >). 
Although Behrens and Henderson (1981) reported finding one specimen off the Palos Verdes 
Peninsula, the other three specimens they listed all came from “Big Kelp Reef, Paradise Cove, Los 
Angeles Co.,” and the geographic coordinates they gave for that site place it in Malibu, not Palos 
Verdes as implied in Behrens and Hermosillo (2005). 
