FIERSTINE ET AL.: CATALOG OF NEOGENE BONY FISHES OF CALIFORNIA 
15 
Referred Specimen in David 09441 : Texas Company Pioneer Unit Plan well number 1, Sunset district, San 
Joaquin Valley, formation not given, Early Miocene, Kern County; scale, whereabouts unknown. 
Distribution of Neogene f L.? zemorrensis outside of California : None known, but it has been reported from 
the Bastendorff Formation, Eocene-Oligocene near Coos Bay, Oregon. 
Remarks : The specimen described in David (1944) is probably the holotype, but since the whereabouts of the 
holotype is unknown, we were unable to check its morphology. 
Order Clupeiformes 
Family Engraulidae 
Genus Engraulis Cuvier 1816 
Engraulis mordax Girard 1854a 
Selected Synonymy & References (fossils only) 
Engraulis mordax Girard: Fitch 1967:9. 
Engraulis mordax Girard 1854a: Fitch and Reimer 1967:79, 81, 84, table 1, fig. 4. 
Engraulis mordax Girard: Fitch 1968:7. 
Engraulis mordax Girard: Fitch 1969a:72, table 1. 
Engraulis mordax Girard: Fitch and Lavenberg 1971:69. 
Engraulis mordax Girard: Nolf 1985:45. 
Engraulis mordax Girard: Grande and Nelson 1985:6 
Type Specimen : Extant, Engraulis mordax Girard, no types known (Eschmeyer 1998:1120). 
Type Locality : San Francisco, California. 
Referred Specimens in Fitch 0967:91 : No specific specimens or locality. He notes that otoliths of E. mordax 
have been found in many Pliocene and Pleistocene deposits in southern California (see Remarks). 
Referred Specimens in Fitch and Reimer 0967:79. 81. 84. table 1. fig. 41 : From LACMIP locality 423 
(= LACM locality 3245), Signal Hill (Interstate 405 and Cherry Avenue, Long Beach), Fernando 
Formation, Plio-Pleistocene, Los Angeles County; one otolith (fig. 4) (See Remarks). 
Referred Specimens in Fitch 0968:71 : No specific specimens or locality. He notes that otoliths have been 
found in many southern California Pliocene and Pleistocene deposits (see Remarks). 
Referred Specimens in Fitch (T969a:table 11 : No specific specimens, but he listed the number of otoliths of 
E. mordax that were found in the following Pliocene localities: 
From LACMIP 471 (= LACM locality 3802), Upper Newport Bay, Upper Fernando Formation, Plio- 
Pleistocene, Orange County; six otoliths. 
From LACMIP 31425, Downtown Los Angeles, Pico Formation, Plio-Pleistocene, Los Angeles County; 
23 otoliths. (See Remarks). 
Referred Specimens in Fitch and Lavenberg 0971:691 : No specific specimens or locality. The authors state 
that otoliths of E. mordax have been found in Pliocene exposures between Mexico and Eureka, California 
(see Remarks). 
Referred Specimens in Nolf ( 1985:45k No specific specimens and refers to the otoliths reported above by Fitch 
(1967). 
Referred Specimens in Grande and Nelson 0985:61 : No specific specimens and refers to the otoliths reported 
above by Fitch (1967). 
Distribution of Neo gene E. mordax outside of California : None known, except the reference above to Mexico. 
Remarks : According to Powell (1998:70-71), LACMIP locality 423 (= LACM locality 31245) is in Pleistocene 
strata, therefore a Pliocene record of E. mordax from this locality is questionable. Fitch (1969a) also list¬ 
ed otoliths of E. mordax from LACMIP locality 435 (Lomita Marl), a locality now considered of 
Pleistocene age (Blake 1991; Dibblee 1999). 
