FIERSTINE ET AL.: CATALOG OF NEOGENE BONY FISHES OF CALIFORNIA 
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Figure 56 t Scopelogadus (?) capistranensis Ebeling 1962, holotype, YPM 3965 (courtesy of Division of Vertebrate Paleontology, 
Peabody Museum of Natural History, Yale University, New Haven, CT, USA); scale bar equals 5 mm. 
Melamphaidae genus and species indeterminate 
Referred Specimens in Fitch and Lavenberg (1968:94) : The authors mention that an “imprint” of a fossil 
melamphaid was found in Miocene strata in Orange County (presumably f Scopelogadus (?) capistra¬ 
nensis), and fossil otoliths of melamphaids have been found in several Miocene, Pliocene, and Pleistocene 
deposits in southern California. 
Order Gasterosteiformes 
Suborder Gasterosteoidei 
Family Gasterosteidae 
Genus Gasterosteus Linnaeus 1758 
Gasterosteus aculeatus Linnaeus 1758 
Selected Synonymy & References (fossils only) 
■fPungitius haynesi David 1945:315, text-fig. 1: Bell 1973a:588. 
Gasterosteus aculeatus Linnaeus 1758:295. 
Gasterosteus aculeatus Linnaeus: Bell 1973a:588, text-fig. 1. 
Gasterosteus aculeatus Linnaeus: Bell 1973b:479-483, text-figs. 1-3. 
Gasterosteus aculeatus Linnaeus: Rawlinson and Bell 1977:586. 
Gasterosteus aculetus aculeatus Linnaeus: Bell 1977:279, text-fig. 1. 
^Gasterosteus abnormis Sytchevskaya and Gretchina 1981:71 -80:Bell 1994:446, table 15.1. 
•f Gasterosteus orientalis Sytchevskaya and Gretchina 1981:71-80: Bell 1994:444, table 15.1. 
Gasterosteus aculeatus Linnaeus: Smith 1981 :table 2. 
Gasterosteus aculeatus Linnaeus: Welton and Link 1982:208, table 2. 
Gasterosteus aculeatus Linnaeus: Bell 1994:440-444, table 15.1. 
Gasterosteus aculeatus Linnaeus: Bell et al. 2009:text fig. IB. 
Type Specimen : Extant, Gasterosteus aculeatus Linnaeus 1758, syntypes (skins) housed in the Zoological 
Society of London (Eschmeyer (1998:34). 
Type locality : Europe. 
Referred Specimens and Holotvoe for iPunzitins h aynesi David : LACM (CIT) locality 389, near Piru Creek, 
Ridge Formation, Middle Pliocene, Ventura County; holotype, LACM (CIT) 10347, right counterpart Bell 
(1973a) also examined a poorly preserved specimen, LACM 28351, from the type locality of -fPungitius 
haynesi. Gasterosteidae. 
Referred Specimens in Bell (1973b) : From LACM locality 1653 (reported by Bell as LAV 1653), Soboda 
Indian Reservation, Bautista Formation, Plio-Pleistocene, Riverside County; LACM 27570 (text-fig. 3a), 
27572-27573,27574 (text-fig. 1), 27575-27585,27586 (text-fig. 2) and UCR 4047/4-4047/21. The assem¬ 
blage consists of four articulated skeletons and associated elements of one or more individuals on 30 slabs, 
20 of which have counterparts. 
