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OCCASIONAL PAPERS OF THE CALIFORNIA ACADEMY OF SCIENCES 
Number 159 
Referred Specimens in Bell 09771 : From LACM (CIT) locality 3897, Lompoc, Late Miocene, Monterey 
Formation, Santa Barbara County; LACM (CIT) 64577, part and counterpart. Gasterosteidae. 
Referred Spcimens and Holotvpe of f G. ahnormis in Svtchevskava and Gretchina 119811 : From Sakhalin 
Island, Agnevo Formation Middle Miocene, Russia: holotype, PIN 3181/101, complete left counterpart. 
Gasterosteidae. 
Referred Spcimens and Holotvpe of f G. orientalis in Svtchevskava and Gretchina (4981) : From Kamchatka, 
Berezov Formation, Middle Miocene, Russia: holotype, PIN 3844/1, complete right counterpart,. 
Gasterosteidae. 
Referred Specimens in Bell et al. 120091 : From LACM locality 1834, Palos Verdes, probably from the Altamira 
Shale Member of the Monterey Formation, Middle Miocene, Los Angeles County; LACM 150177, right 
counterpart, and specimen LACM (CIT) 64577, listed above under Bell (1977). Gasterosteidae. 
Remarks : Bell (1973:588) placed the locality for f Pungitius haynesi in Los Angeles County rather than 
Ventura County as reported by David (1943). Welton and Link (1982:208) stated that LACM (CIT) local¬ 
ity 389 occurs within the Osito Canyon Shale Member of the Peace Valley Formation, Late Miocene, Los 
Angeles County. 
Distribution of Neogene G. aculeatus within California (other than above) : Bell (1994: table 15.1) cited 
Fierstine (misspelled as Fierstein) et al. unpublished data): near Paso Robles, San Luis Obispo County, 
Paso Robles Formation, Plio-Pleistocene. 
Distribution of Neogene G. aculeatus outside of California : Bell (1994: table 15.1) cited the following: Bell 
(unpublished data and 1974) west central Nevada, mid-Late Miocene; Gaudant (1979, 1981) Netherlands, 
Plio-Pleistocene; Sytchevskaya and Gretchina (1981:71-80) Sakhalin Island, Russia, Middle Miocene and 
Kamchatka, Russia, Middle Miocene. 
Gasterosteus cf. G. aculeatus Linneaus 1758 
Referred Specimens of G. cf. G. aculeatus in Casteel and Rvmer 0975: 620-621. Figs. 2B. C : From USGS 
vertebrate locality M-1239, Cache Formation, Plio-Pleistocene, near Lower Lake, Lake County; USNM 
184073, partial right lateral scute (fig. 2B); USNM 184074, partial scute (fig. 2C). 
Gasterosteus sp. 
Selected Synonymy & References 
Gasterosteidae sp. undetermined.: Reynolds 1991:89 
Gasterosteidae sp. undetermined: Bell 1994:table 15.1 
Gasterosteus sp.: Bell 1994:441 
Gasterosteus sp.: Bell and Reynolds 2010:164, 168, fig. 2, appendix 
Referred Specimen in Reynolds 09911 : From SBCM locality 1.76.43, Middle Miocene, Toomey Hills, San 
Bernardino County; pelvic spine. 
Referred Specimens in Bell and Reynolds 12010:164. 168. fig. 2. appendix! : From SBCM locality 1.76.43, 
Argonaut site, Toomey Hills, Middle Miocene, San Bernardino County; SBCM L1814-1012, probable 
stickleback spine; L1814-1013, possible left pelvic or median spine; L1814-1014, left pelvic spine; 
L1814-1015, left pelvic spine; L1814-1016, median spine base; 1814-1017, badly worn spine. 
Suborder Syngnathoidei 
Family Syngnathidae 
Subfamily f Hipposyngnathinae 
Figure 57 
Genus f Hipposyngnathus Danil’chenko 1960 
t Hipposyngnathus imporcitor Fritzsche 1980 
