FIERSTINE ET AL.: CATALOG OF NEOGENE BONY FISHES OF CALIFORNIA 
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Genus Makaira Lacepede 1802 
Makaira nigricans Lacepede 1802 
Selected Synonymy & References 
Makaira sp.: Fierstine and Applegate 1968:32. 
Makaira sp.: Fierstine 1974:37; fig. 9. 
Istiophoridae: Bames et al. 1981:58. 
Makaira : Fierstine and Welton 1988:14, 17. 
Makaira sp.: Fierstine 1990:table 2. 
Makaira sp.: Schultz 1996:22. 
Makaira nigricans Lacepede: Fierstine 1999:435, table 3. 
Makaira nigricans Lacepede: Fierstine 2001:37, table 6. 
Makaira nigricans Lacepede: Camevale et al. 2002:table 3. 
Makaira nigricans Lacepede: Fierstine 2008:45, 48. 
Type Specimen : Extant, Makaira nigricans Lacepede, holotype sketch of a 365 kg adult sent to Lacepede in 
the MNHN (Nakamura, 1983:329). 
Type Locality : lie de Re, Bay of Biscay, France (Nakamura 1983:329). 
Referred Specimens in Fierstine and Applegate (19681 and Fierstine 09741 : From LACM locality 1945, 
Monterey Formation, Late Miocene, Orange County; LACM 17693, rostral fragment. 
Referred Specimens in Bames et al. 09811 and Camevale et al. 120021 : From UCMP locality V68147 
(=LACM locality 4297), San Mateo Formation, latest Miocene, San Diego County; UCMP 118559 
(= LACM 119247), several skull bones and a vertebra. 
Referred Specimens in Fierstine and Welton (1988L Fierstine 0990. 1999L Schultz 09961 : both specimens 
above (LACM 17693 and UCMP 118559). 
Referred Specimens in Fierstine 12008k From OCPC locality 3101, Oso Member, latest Miocene, Capistrano 
Formation, Orange County; both specimens above; OCPC 31001, a nearly complete skull. 
Distribution of Neogene M. ni erica ns andM nit*ricans-\\ke specimens outside of California : From Trinidad 
Formation, Late Miocene- Late Pliocene, Baja California Sur, Mexico: M. nigricans (Fierstine et al. 
2001 ). 
From Yorktown Formation, Early Pliocene, North Carolina, U.S.A.: M. nigricans and Makaira sp. cf. M. 
nigricans (Fierstine 2001). 
From middle or Late Miocene, Anvers, Belgium: ?M. nigricans (= Brachyrhynchus belgicus Leriche 
1926, holotype) (Fierstine 2006). 
From Eastover Formation, Late Miocene, Virginia, U.S.A.: Makaira sp. cf. M. nigricans (Fierstine 1998). 
From Petra Leccese, Late Miocene, Italy: Makaira sp. cf. M. nigricans (Camevale et al. 2002). 
Distribution of Neogene Makaira sd. outside of California : Early Pliocene, near Andalusia, Spain (Schultz 
1996). 
Istiophoridae genus and species undetermined 
Selected Synonymy & References 
Istiophoridae genus and species undetermined:Fierstine and Applegate (1968:30). 
Makaira sp.: Fierstine and Applegate (1968:32). 
Makaira sp.: Fierstine (1974:37). 
Istiophoridae genus and species undetermined: Fierstine (2001:37). 
Istiophoridae genus and species undetermined: Atterholt et al. (2007). 
Referred Specimens of Istiophoridae genus and species undetermined in Fierstine and Applegate 09681 : 
From LACM locality 1625, Sharktooth Hill, Middle Miocene, Kem County; LACM 17695, precaudal ver¬ 
tebra. 
Referred Specimens of Makaira sd. in Fierstine and Applegate 09681 : From LACM locality 65119, Monterey 
Formation, Late Miocene, Orange County; LACM 16074, predentary. 
