FIERSTINE ET AL.: CATALOG OF NEOGENE BONY FISHES OF CALIFORNIA 
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Order Stephanoberyciformes 
Family Melamphaidae 
Melamphaes Gunther 1864 
Melamphaes bispinosus Gilbert 1915 
Synonymy & References 
Melamphaes bispinosus Gilbert 1915:325. 
Scopelogadus bispinosus (Gilbert): Fitch 1968:11. 
Melamphaes bispinosus Gilbert: Eschmeyer 1998:235. 
Type Specimen : Extant, Melamphaes bispinosus Gilbert 1915, holotype, USNM 75809 (Eschmeyer 
1998:235). 
Type Locality : Off Coronado Island, near San Diego, California. 
Referred Specimen in Fitch (1968:11) : He states that otoliths have been found in the Lomita Marl, Pliocene 
(presumably LACMIP locality 435 [= LACM localities 3065, 3759] Miraleste Canyon, Lomita Marl 
Formation, Pleistocene [Irvingtonian], Los Angeles County). 
Remarks : LACMIP locality 435 (Lomita Marl, Los Angeles County) is now considered Pleistocene (Blake 
1991; Dibblee 1999), therefore the record ofM bispinosus in the Pliocene is erroneous. 
Melamphaes lugubris Gilbert 1890 
Selected References 
Melamphaes lugubris Gilbert 1890:59. 
Melamphaes lugubris Gilbert: Fitch 1968:10. 
Type Specimen : Extant, Melamphaes lugubris Gilbert 1890, holotype, USNM 44287 (Eschmeyer 1998:949- 
950). 
Type Locality : California coast south of Point Conception. 
Referred Specimens in Fitch (1968:10) : No specific specimens. Otoliths have been found infrequently in the 
Lomita Marl, Pliocene (presumably LACMIP locality 435 [= LACM localities 3065, 3759], Miraleste 
Canyon, Lomita Marl Formation, Pleistocene [Irvingtonian], Los Angeles County). 
Distribution of Neogene M. lusubris outside of California : None known. 
Remarks : LACMIP locality 435 (Lomita Marl, Los Angeles County) is now considered Pleistocene (Blake 
1991; Dibblee 1999), therefore the record ofM lugubris in the Pliocene is erroneous. 
Order Gasterosteiformes 
Suborder Gasterosteoidei 
Family Gasterosteidae 
Genus Gasterosteus Linnaeus 1758 
f Gasterosteus doryssus (Jordan 1907) 
Selected Synonymy & References 
f Merriamella doryssa Jordan 1907:131, figs. 25-26. 
f Gasterosteus williamsoni leptosomus Hay 1907:271, fig. 12. 
f Gasterosteus doryssus (Jordan): Jordan 1908:117. 
f Gasterosteus doryssus (Jordan): Uyeno and Miller 1963:17, table 1. 
f Gasterosteus doryssus (Jordan): Smith 1981:175, table 2. 
f Gasterosteus doryssus (Jordan): Bell 1994:438-471, table 15.1, figs. 15.2, 15.4, 15.5. 
