MARQUES ET AL.; AMPHIBIANS AND TERRESTRIAL REPTILES OF ANGOLA 
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Global conservation status (lUCN): Least Concern. 
Global distribution: The species is known from southeastern Uganda eastward to coastal 
Kenya and Tanzania, south through Mozambique to KwaZulu-Natal. Also present in Swaziland, 
Botswana, northern Namibia, and Angola. 
Ocurrences in Angola (Map 16): The 
species is distributed across the middle of 
Angola, from the coast to the country’s eastern 
border with Zambia and the Democratic 
Republic of Congo. Kwanza Norte: “Dondo, 
bank of Quanza River” [-9.68333, 14.43333] 
(Poynton and Haacke 1993:13; Ruas 1996:21, 
Ruas 2002:142). Malanje: “Reserva da Palan- 
ca Preta (Rio Caluando)” [-11.46667, 
17.70000] (Ruas 1996:21, 2002:141). Moxico: 
“Lago Dilolo” [-11.50000, 22.01667] (Ruas 
1996:21, 2002:141); “Luso” [-11.78333, 
19.91667] (Ruas 2002:141); “Reserva da 
Palanca Preta (Rio Calombe)” [-11.83333, 
19.93333] (Ruas 1996:21, 2002:141); 
“Luvuei” [-13.06667, 21.16667] (Ruas 
1996:21, 2002:141). Benguela: “Benguella 
(Benguela)” [-12.58333, 13.41667] (Bocage Distribution of Sderophrys gutturalis in 
1867a:845; Loveridge 1936a:82, 1957:310; 
Ruas 1996:21, 2002:142); “Dombe” [-12.95000, 13.10000] (Bocage 1895a:185; Perret 1976a:18). 
Huila: “Sa da Bandeira” [-14.91667, 13.50000] (Ruas 1996:21, 2002:141); “Humpata” 
[-15.03333, 13.40000] (Poynton and Haacke 1993:13; Ruas 1996:21, 2002:142). Namibe: “Mos- 
samedes” [-15.20000, 12.15000] (Bocage 18676:277); “Cima/Saco do Giraul” [-15.06667, 
12.15000] (Poynton and Haacke 1993:13; Ruas 1996:21; 2002:142); “Rio Coroca” [-15.78333, 
12.06667] (Poynton and Haacke 1993:13; Ruas 1996:21, 2002:142). Cuando Cubango: “Cuban¬ 
go basin (2)” [-15.70452, 17.45897] (Conradie et al. 2017:8-9, 12); “Cubango basin (29)” 
[-17.87291, 19.83333] (Conradie et al. 2017:9-10, 12); “Cuito basin (30a)” [-17.50875, 20.06594] 
(Conradie et al. 2017:9-10,12); “Cuito basin (30d)” [-17.51327,20.06111] (Conradie et al. 2017:9- 
10, 12). “Cuito basin (32)” [-17.04880, 19.05333] (Conradie et al. 2017:9-10, 12); “Cuando basin 
(44)” (Conradie et al. 2017:12); “Cuando basin (44a)” [-17.57333, 23.26000] (Conradie et al. 
2017:9-10, 12). 
Taxonomic and distributional notes: The nomen Bufo spinosus Bocage, 1867 was preoccu¬ 
pied by the Iberian Spiny-toad (Daudin 1803) (Loveridge 1936). Prior to this recognition, it had 
been considered a synonym of Bufo regularis Reuss, 1833 by Boulenger (1882) and Bocage 
(1895a). Power (1927) described this taxon as the subspecies Bufo regularis gutturalis based on 
material from the border of Botswana and the Northen Cape. This taxon was later elevated to full 
species by Tandy and Keith (1972), and is currently uniformly accepted as a valid species (Chan- 
ning 2001; du Preez and Carruthers 2009; Frost 2016; Conradie et al. 2016). 
Sderophrys lemairii (Boulenger, 1901) Lemaire’s Toad 
Bufo lemairii-. Boulenger 1901:1, PI. 1, fig. 1. Holotype: not stated, but presumably originally in MRAC (eol- 
leetor C. Lemaire). Type loeality: “Pweto, sur le lae Moero” [= Pweto, Moero Lake], Demoeratie Repub- 
lie of Congo. 
