76 
PROCEEDINGS OF THE CALIFORNIA ACADEMY OF SCIENCES 
Series 4, Volume 65, Supplement II 
Ocurrences in Angola (Map 19): The 
species occurs in western Angola. Malanje: 
“Duque de Braganga falls (Duque de Bragan- 
9a)” [-9.13333, 16.06667] (Poynton and Haac- 
ke 1993:13; Ruas 1996:21, 2002:141). Bie: 
“Caccuchi River” [-13.593333, 16.879861] 
(Poynton et al. 2016:89). Huambo: “Santo 
Amaro” [-12.70000, 15.85000] (Monard 
1937a:26, 1938:77); “Huambo” [-12.737167, 
15.81825] (Poynton et al. 2016:89); “Cubango 
basin (12a)” [-13.59333, 16.87986] (Conradie 
etal. 2016:14). Benguela: “31 kmNE of Sousa 
Lara-Chila (Bocoio)” [-12.46667, 14.13333] 
(Poynton and Haacke 1993:13; Ruas 1996:21); 
“Ebanga” [-12.73333, 14.73333] (Monard 
1937a:26, 1938:78); “Monguavalo Farm” 
[-13.45000, 14.61667] (Poynton and Haacke 
1993:13; Ruas 1996:21). Hufla: “Kalukembe 
(Caluquembe)” [-13.78333, 14.68333] (Monard 1938:55, 76; Ruas 1996:21, 2002:142); “Kuvan- 
gu” [-14.46667, 16.30000] (Monard 1937a:26, 1938:77). Namibe: “Leba pass, between river and 
highway” [-15.07033, 13.24381] (Ceriaco et al. 2016a:19); “Cainde” [-15.48333, 13.36667] 
(Poynton and Haacke 1993:13; Ruas 1996:21; Ceriaco et al. 2016a:19); “16 km W of Vila Nova” 
(Poynton and Haacke 1993:13; Ruas 1996:21; Ceriaco et al. 2016a:19). Cuando Cubango: 
“Cubango basin (3)” [-14.94277, 17.71863] (Conradie et al. 2016:14); “Cubango basin (6a)” 
[-14.67155, 17.73525] (Conradie et al. 2016:14); “Cubango basin (6b)” [-14.67458, 17.73544] 
(Conradie et al. 2016:14); “Cubango basin (20)” [-14.67175, 17.15331] (Conradie et al. 2016:14); 
“Cubango basin (22d)” [-14.64991, 16.90739] (Conradie et al. 2016:14); “Cuito basin (24)” 
[-14.60622, 18.46722] (Conradie et al. 2016:14); “Cubango basin (29)” [-17.87291, 19.83333] 
(Conradie et al. 2016:14); “Cuito basin (30a)” [-17.50875, 20.06594] (Conradie et al. 2016:14); 
“Cuito basin (32)” [-17.04880, 19.53333] (Conradie et al. 2016:14); “Cuando basin (41)” (Conra¬ 
die et al. 2016:14); “Cubango basin (47)” [-14.70214, 17.37806] (Conradie et al. 2016:14); “Cuito 
basin (62)” [-17.50875, 20.06608] (Conradie et al. 2016:14). 
Taxonomic and distributional notes: Tandy and Keith (1972) placed Bufo pusillus {sensu 
Pienaar, 1963) in the synonymy of Sclerophrys maculata (Hallowell, 1854) due the similar mor¬ 
phology and mating calls. While Hulselmans (1969) questioned this synonymy, immunological 
evidence (Maxson 1981; Pickergill 2007a) suggested that it is not distinct from S. maculata. 
A recent study conducted by Poynton et al. (2016) established that S. maculata is now restricted to 
West Africa from the Gulf of Guinea eastwards to Cameroon, while S. pusilla is found in eastern 
and southern Africa, including Angola. Conradie et al. (2016) collected some new material from 
Cuando Cubango Province identified as S. pusilla. Monard (1937a, 1938) identified specimens 
from “Kalukembe” as Bufo funereus (Bocage) [= Sclerophrys funera (Bocage, 1866)] but we 
recently determined these to be S. maculata (Ceriaco et al. 2016b), which now should be refered 
to S. pusilla. 
12°E Ifir'E 20'°:E I4'*E 
Map 19 . Distribution of Sclerophrys pusilla in Angola. 
