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PROCEEDINGS OF THE CALIFORNIA ACADEMY OF SCIENCES 
Series 4, Volume 65, Supplement II 
Ruas 1996:21); “Pungo-Andongo (Pungo Andongo)” [-9.66667, 15.58333] (Boeage 1895a:185; 
Boulenger 1905:107; Ruas 1996:21); “Capanda” [-9.72841, 15.34585] (Ceriaeo et al. 20146:669); 
“Marimba” [-8.36667, 17.03333] (Boulenger 1905:107; Ruas 1996:21) “Cangandala National 
Park” [-9.84606, 16.72233] (Ceriaeo et al. 2016b:27). Mexico: “Teixeira de Sousa, Distr. Villa 
Luso” [-10.70000, 22.23333] (Mertens 1937a: 17); “environs du lae Calundo, rives de la Lumeje, 
Moxieo (Rio Lumege)” [-11.51667, 20.76667] (Laurent 1964a: 130; Ruas 1996:21); “Rives du lae 
Calundo Moxieo” [-11.80000, 20.86667] (Laurent 1964a: 130; Ruas 1996:21); “Reserve de Chasse 
de Cameia, 120 km a Test de Luso (Cameia)” [-11.83333, 21.00000] (Laurent 1964a:130; Ruas 
1996:21); “Cazombo” [-11.88333, 22.91667] (Laurent 1964a:130; Ruas 1996:21). Bie: “Bihe 
(Bie)” [-12.38333, 16.95000] (Boeage 1879b:78, 1895a:185; Ruas 1996:21); “Gauea” [-11.18333, 
17.45000] (Sehmidt 1936:128; Monard 1938:55; Ruas 1996:21); “Chitau” [-11.43333, 17.15000] 
(Sehmidt 1936:128; Monard 1938:55; Ruas 1996:21); “General Maehado” [-12.03333, 17.46667] 
(Mertens 1937a:17). Huambo: “Bimbi (Bimbe)” [-11.81667, 15.83333] (Monard 1937a:26, 
1938:77; Ruas 1996:21); “Cuito, Mt. Moeo” [-12.45000, 15.26667] (Parker 1936:145; Ruas 
1996:21); “Santo Amaro” [-12.70000, 15.85000] (Monard 1937a:26, 1938:77; Ruas 1996:21); 
“Cuma” [-12.86667, 15.06667] (Loveridge 1936a:81). Benguela: “Benguella (Benguela)” 
[-12.58333, 13.41667] (Boeage 1895a:185; Ruas 1996:21); “Dombe” [-12.95000, 13.1000] (Boea¬ 
ge 1895a:185); “Entre Rios” [-13.01667, 14.63333] (Hellmieh 1957a:23); “Cubal” [-13.03333, 
14.25000] (Mertens 1938a:426; Ruas 1996:21); “Cubal da Ganda, Mareo de Canavezes (Mareo de 
Canavezes)” [-13.08333, 14.33333] (Laurent 1964a:130; Ruas 1996:21); “Hanha” [-13.30000, 
14.20000] (Boeage 1896a:113). HmTa: “Caeonda” [-13.73333, 15.06667] (Boeage 1895a:185); 
“Kalukembe (Caluquembe)” [-13.78333, 14.68333] (Monard 1937a:26, 1938:77; Ruas 1996:21); 
“Sangeve (Sangueve)” [-13.88333, 15.83333] (Monard 1937a:26, 1938:77; Ruas 1996:21); “Qui- 
lengues” [-14.10000, 14.08333] (Laurent 1954a:70; Ruas 1996:21); “Kuvangu (Cubango)” 
[-14.46667, 16.30000] (Monard 1937a:26, 1938:77; Ruas 1996:21); “Kapelongo (Capelongo)” 
[-14.88333, 15.08333] (Monard 1937a:26, 1938:77; Ruas 1996:21); “Osi (Osse)” 
[-15.08333, 15.41667] (Monard 1937a:26, 1938:77; Ruas 1996:21); “Kangela (Cangela)” 
[-15.41667, 15.73333] (Monard 1937a:26, 1938:77; Ruas 1996:21). Nambie: “Mossamedes” 
[-15.20000, 12.15000] (Boeage 1895a:185; Ruas 1996:21). Cunene: “Kuvelai (Cuvelai)” 
[-15.65000, 15.80000] (Monard 1937a:26, 1938:77; Ruas 1996:21); “Mupa” [-16.18333, 
15.75000] (Monard 1937a:26,1938:77; Ruas 1996:21). Cuando Cubango: “Kakindo (Caquindo)” 
[-15.45000, 17.05000] (Monard 1937a:26, 1938:77; Ruas 1996:21). Undetermined Locality: 
“from almost all localities that he [J. d’Anchieta] visited” (Boeage 1895a: 185); “Locomi” 
(Boulenger 1905:107; Ruas 1996:21); without precise locality (Laurent 1954:70); “Carangigo” 
(Boulenger 1882:298); “Between Benguela and Bihe” (Boulenger 1905:107). “areas of forest and 
savanna in the north and northeast of Angola” (Cei 1977:16); “arid territories along the coast” (Cei 
1977:17). 
Taxonomic and distributional notes: Sclerophrys regularis (Reuss, 1833) remains a taxo¬ 
nomic challenge. While many specimens from Angola have been identified as S. regularis, Ruas 
(1996) noted that the Angolan records are not representative of Bufo regularis sensu stricto and 
may represent Bufo maculatus (Hallowell, 1854) (currently Sclerophrys pusilla (Mertens, 1937), 
see S. pusilla account) and/or Bufo guturallis (Power, 1927). Sclerophrys regularis is a widely 
distributed in Africa ((lUCN SSC Amphibian Specialist Group 2016), but the boundaries between 
S. regularis and S. gutturalis in Angola, the Democratic Republic of Congo, Kenya, Uganda, and 
Tanzania remain poorly understood. Further studies are needed to clarify the distribution and the 
known range should be regarded as provisional. Ruas (1996) cited several localities (e.g., “Huila,” 
“Bibala,” “Quibula,” “Quindumbo,” “Cahata,” “Dondo,” “Ambaca,” “Catumbela,” “Galanga,” 
