MARQUES ET AL.; AMPHIBIANS AND TERRESTRIAL REPTILES OF ANGOLA 
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undo” [-15.63333, 15.20000] (Monard I937a:30, 1938:56, 83; Ruas 1996:22). Undetermined 
Locality: without precise locality (Bocage 1895a: 181); “Western subregion and Angolan highlands 
(Angolan coastal or watersheds)” (Frade 1963:254); “plateaus regions” (Cei 1977:17); “arid terri¬ 
tories along the coast” (Cei 1977:18). 
Taxonomic and distributional notes: Poynton and Broadley (1985a) regarded this species as 
conspecific with Phrynomantis microps Peters, 1875, which is known from the savannas of West 
and Central Africa, and is now accepted as a valid species. Ruas (1996) suggested that Phryno¬ 
mantis bifasciatus (Smith, 1847) is likely widely distributed in Angola. 
Family Brevicipitidae Bonaparte, 1850 
Genus Breviceps Merrem, 1820 
Breviceps sp. 
Breviceps gibosus Bocage (1870:68). 
Breviceps gibbosus Bocage (1873b:227). 
Breviceps mossambicus: Bocage (1895a:182), Parker (1934:194), Monard (1937a:29, 1938:56, 81), Loveridge 
(1957:357), Inger (1959:532), Laurent (1964a: 156), Cei (1977:17, 18), Gavetti and Andreone (1993:114), 
Ruas (1996:22). 
Rana mossambicus: Hellmich (1957a:30). 
Breviceps mossambicus/adspersus: Poynton (1982:67, 1992:68), Frost (1985:356, 2016), Channing 
(2001:213), du Preez and Carmthers (2009:108), Fretey et al. (2011:27). 
Breviceps adspersus: Poynton and Broadley (1985a:523), Conradie et al. (2016:11). 
Breviceps mossambicus-adspersus: Ruas (2002:142). 
Global distribution: What appears to be a single taxon, though of uncertain identity, ranges 
across west-central Angola. This may belong to a more widely distributed species in southern 
Africa, although phylogenetic support for this is not yet available. 
Ocurrences in Angola (Map 24): 
Records are reported from across western and 
eastern Angola. Lunda Sul: “Alto Chicapa, 
Lunda” [-10.88333, 19.23333] (Laurent 
1964a: 156; Ruas 1996:22, 2002:142); “Riv. 
Luhemba, Alto Chicapa (Rio Luemba)” (Lau¬ 
rent 1964a: 156; Ruas 1996:22, 2002:142). 
Moxico: “Rives du lac Calundo (Lago Calun- 
do)” [-11.80000, 20.86667] (Laurent 
1964a:156; Ruas 1996:22, 2002:142); “Calom- 
be, Luso” [-11.83333, 16.25000] (Ruas 
1996:22; 2002:142); “Cazombo” [-11.88333, 
22.91667] (Laurent 1964a:156; Ruas 1996:22, 
2002:142). Bengo: “R. Donda, 300 miles 
inland from St. Paul de Loanda (Rio Donda)” 
[-9.68333, 14.43333] (Parker 1934:195; Ruas 
1996:22, 2002:142). Huambo: “Bimbi 
(Bimbe)” [-11.81667, 15.83333] (Monard Map 24. Distribution of Sr.,sp. in Angola. 
1937a:29, 1938:81; Monard 1938:56; Ruas 1996:22, 2002:142); “Galanga” [-12.06667, 15.15000] 
(Monard 1938:56; Ruas 1996:22, 2002:142). Benguela: “Quissange” [-12.43333, 14.05000] 
(Bocage 1895a: 182; Parker 1934:195; Monard 1938:56; Ruas 1996:22, 2002:142); “Quindumbo” 
