MARQUES ET AL.; AMPHIBIANS AND TERRESTRIAL REPTILES OF ANGOLA 
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Family Hemisotidae Cope, 1867 
Genus Hemisus Wagler, 1827 
Hemisus guineensis Cope, 1865 Guinea Snout-Burrower 
Hemisus guineensis Cope 1865:100. Holotype: NHMW 1095 (collector unknown) fide Haupl and Tiedemann 
(1978:18). Type locality: Not stated, later restricted to “Guinea,” by inference according to Haupl and 
Tiedemann (1978:18). 
Hemisus guineensis mierops Laurent 1972:55, fig. 27. Holotype: MRAC 851 (collector H. Schouteden). Type 
locality: “Kidada, Lower-Congo,” Democratic Republic of Congo. 
Hemisus guttatum: Bocage (1895a:184), Monard (1938:56), Frade (1963:254). 
Hemisus marmoratum: Monard (1938:56). 
Hemisus marmoratus: Bocage (1887b:208). 
Hemisus marmoratus guineensis: Laurent (1950a: 15), Hellmich (1957a:28), Loveridge (1957:355). 
Hemisus guineensis: Laurent (1964a: 147), Haupl and Tiedemann (1978:18), Channing (2001:121), Rddel and 
Ernst (2003:27), Channing and Howell (2006:126), Onadeko and Rodel (2009:5), Fretey et al. (2011:27), 
Frost (2016), Ceriaco et al. (2016b:29). 
Hemisus guineensis mierops: Cei (1977:17, 18), Poynton and Broadley (1985a:533), Poynton (1992:71), Ruas 
(1996:28), Conradie et al. (2016:11). 
Global conservation status (lUCN): Least Concern. 
Global distribution: The species is known from Angola, south to the Caprivi Strip in Namib¬ 
ia, northern Botswana, and Mozambique to the extreme northeast border of South Africa, and also 
occurs from Senegal to Uganda and Kenya. 
Ocurrences in Angola (Map 25): The 
species is found in western and eastern Angola. 
Zaire: “St. Salvador do Congo” [-6.26667, 
14.23333] (Bocage 1895a;183; Monard 
1938:56; Laurent 1972:55). Kwanza Norte: 
“Mucoso” [-9.53333, 14.65000] (Hellmich 
1957a:28). Malanje: “Cangandala National 
Park” [-9.84606, 16.72233] (Ceriaco et al. 
2016b:29). Lunda Norte: “Muita, Luembe E 
(Muita)” [-7.80000, 21.45000] (Laurent 
1950a: 15; Laurent 1972:55; Ruas 1996:28). 
Moxico: “Cazombo” [-11.88333, 22.91667] 
(Laurent 1964a: 147; Laurent 1972:55; Poynton 
1992:71; Ruas 1996:28). Namibe: “Interieur 
de Mossamedes (Mossamedes)” [-15.20000, 
12.15000] (Bocage 1887b:208; 1895a:184; 
Monard 1938:56; Ruas 1996:28). Undeter¬ 
mined Locality: “arid subregion, Angola” 
(Frade 1963:254); “plateaus regions, Angola” (Cei 1977:17); “arid territories along the coast, 
Angola” (Cei 1977:18). 
Taxonomic and distributional notes: See notes below for Hemisus marmoratus. 
Hemisus marmoratus (Peters, 1854) Marbled Snout-Burrower 
Engystoma marmoratum Peters 1854:628. Syntypes: ZMB 3548 [2 specimens] and ZMB \Q\{)l,fiide Bauer 
et al. (1995:41, 1996:260) (collector W.C.H. Peters). Type locality: “Cabaceira” (Peters 1954:628), 
Mozambique. 
12*E l(i*E 2i>°E 24*E 
Map 25. Distribution of Hemisus guineensis in Angola. 
