MARQUES ET AL.; AMPHIBIANS AND TERRESTRIAL REPTILES OF ANGOLA 31 
senting 23 species, including the type material 
for two species of amphisbaenids - Monopeltis 
okavangensis Monard, 1930 (currently consid¬ 
ered a synonym of Monopeltis anchietae 
(Bocage, 1873)) and Amphisbaena ambuellen- 
sis Monard, 1930 (currently considered a syn¬ 
onym of Zygaspis quadrifrons (Peters, 1862)). 
Encouraged by the results of the first expedi¬ 
tion, Monard, in close contact with the Por¬ 
tuguese colonial administration, organized a 
second expedition to Angola that left Europe in 
March 1932 and lasted until November 1933. 
This second expedition revisited some areas 
surveyed in the first, but also continued further 
north covering more areas in Benguela 
Province as well as in Huambo. The herpeto- 
logical results of the second trip were far 
greater and diverse than those of the first. The 
resulting collection comprised nearly 400 spec¬ 
imens, among which were many new species 
records for the country. It also included several 
new species, both among amphibians — 
Hyperolius cinereus Monard, 1937, Hyperolius 
erythromelanus Monard, 1937 (currently con¬ 
sidered a member of the Hyperolius angolensis 
complex), Rana (Ptychadena) keilingi Monard, 
1937, Rana {Ptychadena) buneli Monard, 1937 (currently considered a synonym of Ptychadena 
bunoderma), Bufo regularis humbensis Monard, 1937 (currently considered a member of the Scle- 
rophrys regularis complex), and Cassionopsis {cmvQntXy Kassina) kuvangensis Monard, 1937 — 
and reptiles — Monopeltis devisi Monard, 1937 (currently considered a synonym of Monopeltis 
anchietae (Bocage, 1873)), Monopeltis grand kuanyamarum Monard, 1937 (currently considered 
a synonym ofDalophia pistillum Boettger, 1895), Mabuya (currently Trachylepis) striata angolen¬ 
sis Monard, 1937 (currently Trachylepis monardi nom. nov. see taxonomic account), and 
Tetradactylus lundensis Monard, 1937 (currently considered a synonym of Tetradactylus ellen- 
bergeri). 
Heinrich Ernst Karl Jordan (1861-1959) from the Tring Museum conducted an expedition to 
South-West Africa (currently Namibia) and Angola in 1934. During this trip, he made an important 
collection of reptiles and amphibians from several areas in Angola, ranging from the forests and 
swamps of Congulu and Quirimbo (Kwanza Sul Province) and the open forests of Mount Moco 
and Catengue (Huambo and Benguela provinces respectively), to the dry, granitic, sandy or lime¬ 
stone areas of Bocoio, Lobito and Morro de Pundo (Benguela Province). This material would be 
later studied by Parker (1936) and lead to the description of one snake and two amphibian taxa. 
From 1937 to 1938, Wilhelm Schack (1909-1959) collected material in Cubal (Benguela Province) 
that was sent to the German herpetologist Robert Mertens (1894-1975) at the Senckenberg Muse¬ 
um in Frankfurt. The collection, containing 439 specimens of 42 different species, was the basis 
for the description of Rhoptropus boultoni benguellensis and Agama planiceps schacki (Mertens 
1938). David Sjolander (1886-1954), curator of Goteborgs Naturhistoriska Museum in Sweden, 
Figure 23. Figure Albert Monard with a speeimen of 
Varanus niloticus during the seeond Swiss Expedition to 
Angola (souree Musee d'histoire naturelle de La Chaux-de- 
Fond). 
