30 
PROCEEDINGS OF THE CALIFORNIA ACADEMY OF SCIENCES 
Series 4, Volume 65, Supplement II 
also partieipated in this expedition to the eenter and southwest of Angola (Malanje, Kwanza Sul, 
Benguela, Huambo, Huila and Namibe provinees; Fig. 22). This expedition yielded a huge eollee- 
tion of amphibians and reptiles, of whieh only the snakes (202 speeimens representing 39 forms, 
three of them new to seienee: Mehelya vernayi, Naja nigricollis nigricincta, and Aspidelaps lubri- 
cus cowlesi) were studied and published on by Charles M. Bogert (1908-1992) in 1940. The 
remaining speeimens remain mostly unstudied although speeimens eolleeted on the Vemay Expe¬ 
dition were reeently used in the deseription of the new Angolan endemie eordylid Cordylus 
namakuiyus Stanley et al., 2016 and by the British/Ameriean zoologist Arthur Loverdige (1891- 
1980) to deseribe Afroedura karroica bogerti (eurrently Afroedura bogerti) and Pachydactylus 
scutatus angolensis (Loverdige 1944), elevated to full speeies by Bauer et al. (2002a). 
Also with the intent of eolleeting speeimens of the Giant Sable for the ereation of a diorama, 
the ANSP organized the Gray Expedition to Angola led by the Ameriean hunter Prentiss N. Gray 
(1884-1935). In the first of the two expeditions (1929), Gray was aeeompanied by the ornitholo¬ 
gist Wilfrid Wedgwood Bowen (1899-1987) and by an employee of the Companhia de Diamantes 
de Angola, J. R. Evans (birth and death dates unknown). In August 1930, Harold T. Green (1896- 
1967) led a seeond expedition promoted by the ANSP to Angola to eolleet additional material for 
the diorama, and some reptiles and amphibians were eolleeted. This latter expedition took three 
months, beginning in Lobito and then spending most of the time in the area of Malanje, but also 
with a small ineursion into Katanga (then in the Belgian Congo). This small eolleetion, still extant 
in the ANSP, was never published upon. 
From 1930 to 1931, Ralph Pulitzer (1879-1939) and his wife Margaret Kemoehan (nee Leeeh) 
Pulitzer and (Wolfrid) Rudyerd Boulton, Jr. (1901-1983) and his wife Laura Theresa (nee Crayton) 
Boulton (1899-1980) earried out the Pulitzer Angola Expedition that was promoted by the 
Carnegie Museum (CM). This expedition mainly foeused on mammals and birds and explored 
areas in the eenter (Bie Provinee) and southwest (Benguela, Namibe, Huila and Cunene provinees). 
It yielded important herpetologieal results that were published by Karl Patterson Sehmidt (1890- 
1957) at the Field Museum of Natural History (FMNH), Chieago. Due to the involvement of 
Sehmidt, duplieates of some speeies were deposited in the FMNH, while the majority of the mate¬ 
rial remained in CM. Sehmidt published the results in two prineipal papers, the first dedieated to 
reptiles (Sehmidt 1933) and the seeond to amphibians (Sehmidt 1936). The reptile eolleetion, eom- 
prising 457 speeimens belonging to 40 different speeies, was used by Sehmidt (1933) to deseribe 
four new taxa - Rhoptropus boultoni, Pachydactylus bibroniipulitzerae, Lygodactylus laurae, and 
Varanus albigularis angolensis. The amphibian eolleetion ineluded 442 speeimens representing 17 
different speeies, whieh, though not ineluding any new taxa, did represent several important range 
extensions (Sehmidt 1936). 
At about the same time that North Ameriean institutions undertook expeditions in Angola, 
several European institutions and individuals also promoted expeditions to the Portuguese eolony. 
One of the most important eolleetors was the Swiss naturalist Albert Monard (1886-1952; Fig. 23). 
He undertook two large expeditions to Angola for the Museum of La Chaux-de-Fonds (where the 
eolleetions still exist) in eollaboration with the former National Museum of Lisbon, then renamed 
the MuseuBoeage (Monard 1931, 1932, 1937a,b, 1938). 
The first of Monard’s trips was originally designed as a private hunting endeavor organized by 
other Swiss nationals. This trip lasted from July 1928 to February 1929 (but only three months 
were effeetively dedieated to speeimen eolleeting) and explored Chimporo, Ebanga, Kakindo, 
Kuvangu, Kalukembe, Catumbela, Santo Amaro, Vila da Ponte, Bimbi, Caquindo, Chimporo, 
Kuvelai and Mbale, all in the southern areas of the country (Benguela, Huila, Cunene, Bie, Cuan- 
do Cubango provinces). The resulting collection comprised 45 herpetologieal specimens, repre- 
