MARQUES ET AL.: AMPHIBIANS AND TERRESTRIAL REPTILES OF ANGOLA 
27 
Figure 20. Depiction of Anchieta’s laboratory in Caconda (source Arquivo Historico Museu Bocage, artist and date 
unknown). 
from Namibe Province, obtained during the exploration of the Curoea River basin on the second 
(Bocage 1887b). 
Boeage dedicated the following decades to the study of Angolan fauna with a special interest 
in herpetology and ornithology, and he published 24 eontributions on the Angolan herpetofauna 
(Bocage 1864, 1866a,b, 1867a,b,c,d, 1870, 1872, 1873a,b, 1876, 1879a,b,e, 1882a, 1886a, 
1887a,b,e, 1888, 1889, 1890, 1893, 1895a,b, 1896a,b, 1897a,b). As his major eontribution to the 
study of the Angolan herpetofauna, Bocage published “Herpetologie d’Angola et du Congo,” the 
first, and until now, the only eomplete summary of Angolan amphibians and reptiles, eomprising 
at the time a total of 191 speeies (41 amphibians and 150 reptiles; Boeage 1895a). Since the time 
of Bocage, the number of known herpetologieal taxa has more than doubled. 
Under the direet supervision of Boeage, the Portuguese naturalist Jose Julio Betteneourt 
Ferreira (1866-1948) was appointed as assistant in the herpetologieal colleetions of the National 
Museum of Lisbon and started his career studying the Angolan herpetofauna (Ferreira 1897a), 
including the specimens of the last shipment sent by Anehieta (Ferreira 1897b). Bocage died in 
1907, and although his last papers were published in 1905, his age and delieate health effeetively 
ineapaeitated him during the first years of the twentieth eentury. Due to Bocage’s eondition, Bet¬ 
teneourt Ferreira pursued the study of the Lisbon Museum Angolan material, espeeially that sent 
by private collectors and donors (Ferreira 1900, 1903). Ferreira was also responsible for the study 
of the herpetologieal material of Franciseo Newton’s expedition to Angola. Newton (1864-1909; 
Fig. 21), who had colleeted extensively for the Lisbon Museum and Bocage in the Cape Verde 
Islands, the Gulf of Guinea, and Timor, was hired by the Polytechnie Aeademy of Porto to lead an 
expedition to Angola. This expedition (1903-1905) foeused on Luanda, Bengo, Kwanza Norte, 
Kwanza Sul, Malanje, and Namibe provinces. Ferreira published upon the eollections made from 
