84 
PROCEEDINGS OF THE CALIFORNIA ACADEMY OF SCIENCES 
Series 4, Volume 65, Supplement II 
Hemisus marmoratus: Inger (1959:541), Schmidt and Inger (1959:170), Channing (2001:124), Rodel and 
Ernst (2003:27), Onadeko and Rodel (2009:5), Fretey et al. (2011:27), Frost (2016). 
Hemisus marmoratum: Bocage (1887a:183, 1895a:183), Boulenger (1905:107), Monard (1937a:29, 1938:56, 
82), Frade (1963:254). 
Hemisus sudanense: Boulenger (1882:178). 
Hemisus marmoratus (?) angolensis: Hellmieh (1957a:28). 
Hemisus guineensis mierops: Laurent (1972:55), Ruas (1996:28). 
Global conservation status (lUCN): Least Concern. 
Global distribution: The species is widespread in sub-saharan Africa (excluding rainforest 
areas), extending from Senegal to Ethiopia and Somalia, south into northern South Africa. 
Ocurrences in Angola (Map 26): The 
species is found in western and eastern Angola. 
Kwanza Norte: “Dondo” [-9.68333, 
14.43333] (Bocage 1895a:183; Monard 
1938:56; Laurent 1972:55; Ruas 1996:28). 
Kwanza Sul: “Semba Acendu” [-9.85000, 
16.15000] (Boulenger 1905:107; Monard 
1938:56; Laurent 1972:55; Ruas 1996:28). 
Bie: “Chitau” [-11.43333, 17.15000] (Laurent 
1972:55; Poynton 1992:71). Benguela: 
“Catumbella (Catumbela)” [-12.43333, 
13.55000] (Bocage 1895a:183; Monard 
1938:56; Laurent 1972:55; Ruas 1996:28); 
“Benguella (Benguela)” [-12.58333,13.41667] 
(Boulenger 1882:179; Laurent 1972:55, Poyn¬ 
ton 1992:71). Huila: “Kalukembe (Caluquem- 
be)” [-13.78333, 14.68333] (Monard 1937a:29, 
1938:56, 82; Laurent 1972:55; Ruas 1996:28). 
Map 26. Distribution of Hemisus marmoratus in Angola. 
Cuando Cubango: “Kakindo (Caquindo)” [-15.45000,17.05000] (Monard 1937a:29,1938:56, 82; 
Laurent 1972:55; Ruas 1996:28). Undetermined Locality: “western subregion and Angolan high¬ 
lands (Angolan coastal or watersheds)” (Cei 1977:18). 
Taxonomic and distributional notes: The delimitation of Hemisus guineensis (Cope, 1865) 
and Hemisus marmoratus (Peters, 1854) is more complicated than reflected by the current taxono¬ 
my. Both are likely part of a larger complex of morphologically similar species (Rodel and Ernst 
2003; Onadeko and Rodel 2009). Both species have been reported to occur in Angola. However, 
Laurent (1972), in the only synoptic taxonomic review of the genus, explicitly considered Angolan 
populations to represent what he described as a subspecies of H. guineensis {H. guineensis 
mierops) that might even represent a distinct species. In West Africa, H. guineensis occurs in the 
forest or near to forests, and is common in grasslands and open woodlands, whereas H. marmora¬ 
tus is usually associated with savannas (Channing 2001; Channing and Broadley 2002; Rodel and 
Ernst 2003; Channing and Howell 2006; Onadeko and Rodel 2009; Frost 2016). Channing (2001) 
provided several records for both species in Angola but without information about the specimens 
corresponding to these records. Vouchers sampled from diverse habitat types and with associated 
genetic data are needed to evaluate which taxa are present in Angola as well as their distributions. 
