PROCEEDINGS OF THE CALIFORNIA ACADEMY OF SCIENCES 
Series 4, Volume 65, Supplement II 
al. (2005), where it had been placed by Laurent (1982) and recently placed again by Fretey et al. 
(2011). Pickersgill (2007b) suggested that the validity of this species is questionable, which leaves 
the Angolan records for this species in need of further study. 
Afrixalus wittei (Laurent, 1941) de Witte’s Spiny Reed Frog 
Megalixus wittei Laurent 1941:127. Holotype: MRAC 11500. Type loeality: “Lukafu” (Laurent 1941:127), 
Demoeratie Republie of Congo. 
Hyperolius fulvovittatus: Boeage (1866a:55). 
Rappia fulvovittata: Boeage (1895a: 175). 
Afrixalus wittei: Ferret (1976b:24), Frost (1985:224, 2016), Poynton and Broadley (1987:191), Sehiotz 
(1999:64), Channing (2001:143), Fretey et al. (2011:27), Ceriaeo et al. (2016b:30). 
Global conservation status (lUCN): Least Concern. 
Global distribution: The species is known from tropical lowland savannas from the southern 
Democratic Republic of Congo, northeastern Angola and Zambia. 
Ocurrences in Angola (Map 31): The 
species was from some time only known from 
a single record from “Duque de Braganga” 
[= Calandula], Malanje Province. It was recent¬ 
ly found in Cangandala National Park, in the 
same region. Malanje: “Duque de Bragan^a” 
[-9.10000, 15.95000] (Boeage 1866a;55, 
1895a: 175; Ferret 1976b:24); “Cangandala 
National Park” [-9.84606, 16.72233] (Ceriaeo 
etal. 2016b:30). 
Taxonomic and distributional notes: 
Ferret (1976b) considered the specimen from 
“Duque de Braganga” identified by Boeage as 
Hyperolius fulvovittatus (Boeage 1866a), and 
later as Rappia fulvovittata (Boeage 1895a), to 
be referable to Afrixalus wittei (Laurent, 1941). 
Through analysis of the morphological diversi¬ 
ty in several species of Afrixalus, Laurent 31. Distribution of Afrixalus wittei in Angola. 
(1982) argued that A. wittei is closely related to A. osorioi Ferreira, 1906 (Poynton and Broadley 
1987). 
Genus Cryptothylax Laurent and Combaz, 1950 
Cryptothylax greshoffii (Schilthuis, 1889) Greshoee’s Wax Frog 
Hylambates Greshoffii Schilthuis 1889:286, unnumbered fig. Type: UZ by original designation, now possibly 
deposited in RMNH fide Hoogomoed in Frost (1985:205) (eolleetor A. GreshofiE). Type loeality: “Boma 
(Congo, W. Afriea)” (Sehilthuis 1889:284), [= Boma], Demoeratie Republie of Congo. 
Hylambates greshoffii: Frost (1985:205). 
Cryptothylax greshoffii: Sehiotz (1999:90), Blaekbum and Jaekson (2006:358), Fretey et al. (2011:29), Chan¬ 
ning et al. (2012:200), Frost (2016). 
Global conservation status (lUCN): Least Concern. 
Global distribution: The species is known from the forests of southern Cameroon extending 
into the Democratic Republic of Congo, and south to northern Angola. 
Ocurrences in Angola: The species occurs in northern Angola. Uige: without precise location 
(Sehiotz 1999:91). 
