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PROCEEDINGS OF THE CALIFORNIA ACADEMY OF SCIENCES 
Series 4, Volume 65, Supplement II 
lensis (Bocage, 1893) (Channing et al. 2013b; Ceriaco et al. 2014a; see H. benguellensis and 
H. nasutus accounts). The syntypes of R. nobrei are similar to both H. adspersus and H. benguel¬ 
lensis, which makes it difficult to determine to which species Rappia nobrei should be referred. We 
here follow Ceriaco et al. (2014a) by considering R. nobrei to be a junior synonym of H. adsper¬ 
sus. Channing et al. (2013b) comidQrQd Hyperolius granulatus (Boulenger, 1901), previously asso¬ 
ciated with Hyperolius benguellensis (Bocage, 1893), to be a synonym of H. adspersus (Ferret 
1976a; Schiotz and Van Daele 2003). 
Hyperolius angolensis Steindachner, 1867 Angolan Reed Frog 
[Note: Because of the complexity of the taxonomy of this species we provide additional commentary on its synonymy/chresonymy] 
Hyperolius marmoratus Rapp 1842:289, pi. 6, figs. 1-2. Type: Not designated (eolleetor unknown). Type 
loeality: “Natal” [= KwaZulu-Natal], South Afriea. 
Hyperolius parallelus Gunther 1858a:326. Syntypes: BMNH 1947.2.9.41-43 [3 speeimens] (eolleetor 
unknown, probably “Mr. Rieh”). Speeimens BMNH 1947.2.9.41 and 1947.2.9.42 are depleted in Gun¬ 
ther “1858b” 1859: 86, pi. 8, fig. A. Type loeality: “Sud-Afrika” later ehanged to “South Afriea” and 
“Angola” (Gunther “1858b” 1859:86) and to “Cape of Good Hope” and “Ambris, Angola” [= Ambriz] 
by Boulenger (1882:121). Synonymized with H. marmoratus by Boulenger (1882:121) and treated as a 
subspeeies by Laurent (1951:38). 
Hyperolius marmoratus var. angolensis Steindaehner 1867:50. Syntypes: NHMW 13487.1-2 fide Haupl and 
Tiedemann (1978:22) (eolleetor F.A.P. Bayao). Type locality: “Angola” (Steindachner 1867:50). 
Restricted to “Duque de Bragan^a” [= Calandula] (Ceriaco et al. 2014b:669). 
Hyperolius insignis Bocage 1867a:844, fig. 2. Syntypes: MBL T.21-164, 27-167 [2 specimens] (collector 
J. A. d’Anchieta [Benguella] and P. Barroso [St. Salvador du Congo]), destroyed by fire 18 March 1978. 
Type locality: “Benguella,” Angola. Corrected by Ferret (1976a: 28) to “St. Salvador du Congo” 
[= M’Banza Congo] and “Novo Redondo” [= Sumbe], Angola. Synonymized with H. marmoratus by 
Boulenger (1882:121) and Ferret (1976a:27) and treated as a subspecies by Laurent (1951 “1952”:39). 
Synonymized with H. parallelus by Loveridge (1953a:329). 
Hyperolius Toulsonii Bocage 1867a:845, fig. 3. Holotype: MBL T.27-275 (collector M. Toulson), destroyed 
by fire 18 March 1978. Type locality: “Loanda” [= Luanda], Angola. Synonymized with H. marmoratus 
by Boulenger (1882: 121), recognized as valid by Ahl (1931b:388), and considered a synonym ofH. m. 
insignis by Laurent (1952 “1951”:391), and later synonymized with H. parallelus (Ferret 1976a:27). 
Hyperolius huillensis Bocage 1873b:225. Syntypes: MBL 2-166 [4 specimens] (collector J. A. d’Anchieta), 
destroyed by fire 18 March 1978. Type locality: “Huilla,” [Angola] (Bocage 1873b:225). Considered a 
subspecies ofH. marmoratus by Laurent (1952 “1951”:390). 
Hyperolius vermieulatus Peters 1882a:8. Lectotype: ZMB 10050, designated by Laurent (1961:88) (collector 
von F.W.A. von Mechow). Type locality: “Malange” [=Malanje] (Peters 1882a:8). Considered a sub¬ 
species ofH. marmoratus by Laurent (1952 “1951”:88). 
Rappia plieifera Bocage 1893:118. Syntypes: MBL T.22-209 [2 specimens] (collector J. A. d’Anchieta 
[Caconda] and F. A. P. Bayao [Duque de Braganga]), destroyed by fire 18 March 1978. Haupl and Tiede¬ 
mann (1978:29) considered NHMW 22895 [2 specimens] as part of the type series. Type locality: 
“Caconda,” “Duque de Braganga,” Angola (Bocage 1893:118). Synonymized with/£ m. vermieulatus hy 
Laurent (1952 “1951”:390) and later with H. parallelus by Ferret (1976a:27). 
Rappia marmorata var. marginata: Bocage 1895a: 164: Type: MBL, specimen number(s) not known (collec¬ 
tor F. A. P. Bayao), destroyed by fire 18 March 1978. Type locality: “Duque de Braganga” [= Calandu¬ 
la], Angola. 
Rappia marmorata var. taeniolata Bocage 1895a: 164: Syntypes: MBL, specimen numbers not kn own (col¬ 
lector F. A. P. Bayao [Duque de Braganga], Graga and J. A. d’Anchieta [Huilla] and J. A. d’Anchieta 
[Caconda and Cahata]), destroyed by fire 18 March 1978. Type locality: “Duque de Braganga,” “Huil¬ 
la,” “Caconda” and “Cahata” [= Calandula, Huila, Caconda and Caota], Angola. 
Rappia marmorata var. variegata Bocage 1895a: 164: Syntypes: MBL (collector J. A. d’Anchieta). Type local- 
