MARQUES ET AL.: AMPHIBIANS AND TERRESTRIAL REPTILES OF ANGOLA 
95 
Katanga Province), southern Angola and adja¬ 
cent northern Nambia and Botswana. 
Ocurrences in Angola (Map 34): The 
species is known from western Angola from the 
type locality “Cahata, Benguella,” although 
there are some records further north. Kwanza 
Norte: “Quilombo” [-9.33333, 14.90000] (Fer¬ 
reira 1906:161). Huambo: “Bimbi” 
[-11.81667, 15.83333] (Monard 1938:87). Bie: 
“Cubango basin (11)” [-13.69413, 17.06177] 
(Conradie et al. 2016:8-9, 15); “Cubango basin 
(12a)” [-16.59333, 16.87986] (Conradie et al. 
2016:8-9, 15); Benguela: “Cahata (Caota)” 
[-12.35000, 14.81667] (Boeage 1893:119 
1895a:169, 1897a:204; Perret 1976a:27; Frost 
1985:207, 2016; Poynton and Broadley 
1987:208; Channing et al. 2002:96; Amiet 
2005:275; Channing et al. 2013:318); “Eban- 
ga” [-12.73333, 14.73333] (Monard 1938:87); 
“Hanha” [-13.30000, 14.20000] (Boeage 1897b:211). Huila: “Indungu” [-14.81667, 16.26667] 
(Monard 1938:87); “Zootecnica Plateau, Humpata” [-14.96581, 13.34458] (Channing et al. 
2013:317); “Humpata” [-14.23814, 13.43331] (Channing et al. 2013b:317); “Humpata” 
[-15.03333, 13.40000] (Conradie et al. 2012a:2). Cunene: “Chimporo” [-16.03333, 17.15000] 
(Monard 1937a:39, 1938:94). Cuando Cubango: “Cubango basin (5)” [-14.74628, 17.66844] 
(Conradie et al. 2016:8-9, 15); “Cubango basin (6b)” [-14.67458, 17.73544] (Conradie et al. 
2016:8-9, 15); “ Cubango basin (7b)” [-14.43377, 17.82957] (Conradie et al. 2016:8-9, 15); 
“Cubango basin (7c)” [-14.43916, 17.81491] (Conradie et al. 2016:8-9, 15); “Cubango basin (8)” 
[-14.25705, 17.77852] (Conradie et al. 2016:8-9,15); “Cubango basin (21)” [-14.66586, 17.07661] 
(Conradie et al. 2016:8-9, 15); “Cubango basin (22b)” [-14.66278, 16.96081] (Conradie et al. 
2016:8-9, 15); “Cuito basin (27)” [-15.17127, 19.19433] (Conradie et al. 2016:8-9, 15); “Cuito 
basin (30a)” [-17.50875, 20.06594] (Conradie et al. 2016:9-10, 15); “Cuito basin (30d)” 
[-17.51327, 20.06111] (Conradie et al. 2016:9-10, 15); “Cuito basin (32)” [-17.04880, 19.53333] 
(Conradie et al. 2016:9-10, 15); “Cuito basin (35)” [-16.62322, 19.05352] (Conradie et al. 2016: 
9-10, 15); “Cuando basin (43)” [-17.53500, 23.18916] (Conradie et al. 2016:9-10, 15); “Cubango 
basin (45)” [-16.88350, 18.01180] (Conradie et al. 2016:9, 12, 15). Undetermined Locality: 
“Western subregion and Angolan highlands (Angolan coastal or watersheds)” (Frade 1963:254). 
Taxonomic and distributional notes: In the description, Boeage notes that the type speci¬ 
mens are similar to Rappia nasuta (Gunther, 1865). In their detailed diseussion of the benguelen- 
sis-nasutus complex, Poynton and Broadley (1987) had little doubt that the characters used at that 
time were inadequate to separate all available specimens into nasutus and benguellensis. Schiotz 
(1999) concluded that it is difficult to separate H. nasutus and H. benguellensis based on mor¬ 
phology and color patterns. Channing et al. (2002) proposed formal ehanges in nomenelature and 
gave detailed lists of synonyms, but Schiotz (2006) questioned several of these changes. Schiotz 
and Van Daele (2003) provided a discussion about the differenees in the advertisement calls of 
nasutus and benguellensis as well as differences in external morphology. Hyperolius benguellensis 
(Boeage, 1893) was questionably removed from the synonymy of H. nasutus Gunther, 1865 by 
Amiet (2005). Channing et al. (2013) used molecular data to argue that H. benguellensis is a valid 
12°E \e>E 20°E 
Map 34. Distribution of Hyperolius benguellensis in 
Angola. 
