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PROCEEDINGS OF THE CALIFORNIA ACADEMY OF SCIENCES 
Series 4, Volume 65, Supplement II 
species. The species has only been confirmed from southern Angola, northern Namibia, and north¬ 
ern Botswana where it was found in open grassy habitats, along streams or man-made structures 
with emergent vegetation (Channing et al. 2013). We follow Poynton and Broadley (1987), Schiotz 
and Daele (2003), and Channing et al. (2013) in regarding H. benguellensis as distinct from the 
sympatric H. nasutus. Hyperolius oxyrhynchus is regarded as a synonym of H. benguellensis 
(Channing et al. 2013b), though a record from “Muita” (Laurent 1950) is probably a misidentifi- 
cation as it is far to the north of the rest of the distribution of H. benguellensis. Laurent’s (1950a) 
record should probably be referred to another species of the nasutus complex, such as H. adsper- 
sus or H. nasutus. 
Hyperolius bicolor Ahl, 1931 Two-Colored Reed Frog (Endemic) 
Hyperolius bicolor Ahl 193 la: 129. Syntypes: ZMB, lost (collector K. May). Type locality: “Farenda Bango, 
Loanda” (Ahl 193la: 129), [= Fazenda Bango, Luanda], Angola. 
Hyperolius bicolor. Frost (1985:208, 2016). 
Global conservation status (lUCN): 
Data Deficient. 
Global distribution: The species is 
known only from Angola. 
Ocurrences in Angola (Map 35): The 
species is known only from the type locality 
“Fazenda Bango, Loanda”. Luanda; “Farenda 
Bango” [-9.81767, 14.78916] (Ahl 1931a:129; 
Frost 1985:208, 2016). 
Taxonomic and distributional notes: 
According to Laurent {in Frost 1985:208, 
2016) the species is probably a synonym of 
Hyperolius marmoratus insignis, a member of 
parallelus-marginatus subgroup (Schiotz 
1999) [see Hyperolius angolensis account]. 
The validity of this taxon is questionable and 
difficult to address due to loss of the type spec¬ 
imen. 
12*E I6®E 20°E 24*E 
Map 35. Distribution of Hyperolius bicolor in Angola. 
Hyperolius bocagei Steindachner, 1867 Bocage’s Reed Frog 
Hyperolius bocagei Steidachner 1867:51, pi. 5, fig. 11. Holotype: NHMW 14846 fide Haupl and Tiedemann 
(1978:22) (Collector F.P. Bayao). Type locality: “Angola” (Steindachner 1897:51). 
Rappia bocagei var. maculata Ferreira 1906:160. Type: Not stated (collector F. Newton), see Ceriaco et al. 
(2014a:25-26) discussion. Type locality: “Golungo Alto”, Angola. 
Rappia seabrai Ferreira 1906:163. Holotype: (collector F. Newton), not located, probably lost. Paratype: 
MNHFCP 018587 (collector F. A. P. Bayao). Type locality: “Quilombo, Rio Luinha” [= Gonguembo] 
Kwanza Norte, Angola. 
Hyperolius bocagei: Bocage (1873b:225), Noble (1924:252), Parker (1936:143), Laurent (1950a: 16, 
1954a:79, 1964a: 150), Cei (1977:17); Poynton and Broadley (1987:212), Haupl and Tiedemann (1978:22), 
Frost (1985:208,2016), Haupl et al. (1994:26), Schiotz (1999:188), Channing (2001:151), Schiotz and Van 
Daele (2003:145), Ceriaco et al. (2014a:24). 
Rappia bocagii: Boulenger (1882:126, 1905:109), Bocage (1895a: 165, 1897a:203). 
Rappia cinctiventris: Bocage (1895a: 168), Boulenger (1905:110). 
Rappia bocagei: Ferreira (1904:112, 1906:160). 
