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PROCEEDINGS OF THE CALIFORNIA ACADEMY OF SCIENCES 
Series 4, Volume 65, Supplement II 
Province, is approximately 140 km west of Caladula Falls in Malanje Province, formerly “Duque 
de Branganga,” the type locality of H. bocagei and the locality from which the paratype of 
R. seabrai was collected (Ceriaco et al. 2014a). Schiotz (1999) and Schiotz and Van Daele (2003) 
suggested that H. bocagei might be a junior synonym of a species in the Hyperolius viridiflavus 
complex. Poynton (1986) considered Hyperolius cinctiventris Cope, 1862 to be a synonym of 
Hyperolius argus Peters, 1854, and as this species is restricted to East Africa it is unlikely to 
correspond to Angolan specimens. The specimens identified by Monard (1938) as cinctiventris 
deposited in the Musee d’Histoire Naturelle, La-Chaux-de Fond, Switzerland were recently stud¬ 
ied and identificated as Hyperolius bocagei (Ceriaco et al. in prep.). 
Hyperolius chelaensis Conradie, Branch, Measy and Tolley, 2012 Chela Mountain 
Reed Frog (Endemic) 
Hyperolius chelaensis Conradie et al. 2012a:5, figs 5-6. Holotype: PEM A9223 (eolleetor W. Conradie). Type 
loeality: “a small pateh (< 2.5ha.) of Afromontane forest in a small gorge draining from Serra da Chela 
above the Estaeao Zooteeniea, near Humpata, Lubango” (Conradie et al. 2012a: 5), Angola. 
Hyperolius chelaensis: Conradie et al. (2013:203), Frost (2016). 
Global conservation status (lUCN): Data Deficient. 
Global distribution: The species is known only from Angola. 
Ocurrences in Angola (Map 37): The 
species is known only from the type locality 
“Serra de Chela,” Lubango. Huila: “Serra de 
Chela gorge, above the Estagao Zooteeniea 
near Humapata, Lubango” [-14.88944, 
13.27417] (Conradie et al. 2012a:5; Conradie 
et al. 2013:203). 
Taxonomic and distributional notes: 
This species was recently described by Con¬ 
radie et al. (2012a). According to the original 
publication, H. chelaensis is the sister species 
of H. cinereus Monard, 1937. The type locality 
is along a small stream that drains from the 
Serra de Chela escarpment into the ephemeral 
water courses (e.g., Giraul River) draining west 
to the Atlantic Ocean (Conradie et al. 2013). It 
is expected to be more widespread in the Serra 
de Chela and Leba mountain ranges. Further Map 37. Distribution of Hyperolius chelaensis in 
surveys are needed to establish a full distribu- 
tion of the species and its conservation status. 
Hyperolius cinereus Monard, 1937 Ashy Reed Frog (Endemic) 
Hyperolius cinereus Monard 1937a:32. Syntypes: MHNC 90.0856-7 (collector A. Monard). Type locality: 
“Kalukembe” and “Bimbi” (Monard 1937a:32), [= Caluquembe and Bimbe], Angola. 
Hyperolius cinereus: Monard (1938:85), Cei (1977:17), Frost (1985:209, 2016), Schiotz (1999:192), Chan- 
ning (2001:152), Conradie et al. (2012a:17), Conradie et al. (2013:222). 
Hyperolius {Hyperolius) cinereus: Fretey et al. (2011:30). 
Global conservation status (lUCN): Least Concern. 
