MARQUES ET AL.: AMPHIBIANS AND TERRESTRIAL REPTILES OF ANGOLA 
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Global distribution: The species is 
known only from Angola. 
Ocurrences in Angola (Map 38): The 
species is known from southwestern and cen¬ 
tral Angola. Bie: “western dambo of Cacuchi 
River, Bie Province” [-13.58333, 16.86667] 
(Conradie et al. 2013:222); “Cubango basin 
(11)” [-13.69413, 17.06177] (Conradie et al. 
2016:8-9, 16). Huambo: “Bimbi” [-11.81667, 
15.83333] (Monard 1937a:32, 1938:85, Chan- 
ning 2001:153; Conradie et al. 2013:222); 
“small stream at Huambo Agriculture Institute, 
Huambo Province” [-12.71667, 15.80000] 
(Conradie et al. 2013:222); “1st stream cross¬ 
ing east of Caccuchi River, Bie Province” 
[-13.68333, 17.05000] (Conradie et al. 
2013:222). Benguela: “Zamba-Seidlung bei 
Entre Rois, Benguella Province” [-13.01667, 
14.63333] (Conradie et al. 2013:222). Huila: “Kalukembe” [-13.78333, 14.68333] (Monard 
1937a:32, 1938:85; Frost 1985:209, 2016; Channing 2001:153; Conradie et al. 2013:201, 222); 
“stream under road before Estacao Zootecnica near Humpata, Lubango” [-14.90400, 13.32556] 
(Conradie et al. 2012a: 17; Conradie et al. 2013:222); “waterfall below dams, on plateau above 
Estacao Zootecnica near Humpata, Lubango” [-14.91425, 13.31386] (Conradie et al. 2012a: 17, 
Conradie et al. 2013:222). Cuando Cubango: “5 km south of Soba Matios Military base, Cuan- 
do” [-14.25706, 17.77861] (Conradie et al. 2013:222); “small stream after Muvange river, near 
Menongue” [-14.73333, 17.66667] (Conradie et al. 2013:222); “Muvange river crossing, near 
Menongue” [-14.81667, 17.66667] (Conradie et al. 2013:222); “Cubango basin (5)” 
[-14.74628,17.66844] (Conradie et al. 2016:8-9,16); “Cubango basin (6b)” [-14.67458, 17.73544] 
(Conradie et al. 2016:8-9, 16); “Cubango basin (7c)” [-14.43916, 17.81491] (Conradie et al. 
2016:8-9, 16); “Cubango basin (8)” [-14.25705,17.77852] (Conradie et al. 2016:8-9, 16); “Cuban¬ 
go basin (21)” [-14.66586, 17.07661] (Conradie et al. 2016:8-9, 16); “Cubango basin (22b)” 
[-14.66278, 16.96081] (Conradie et al. 2016:8-9, 16); “Cuito basin (27)” [-15.17127, 19.19433] 
(Conradie et al. 2016:8-9, 16); “Cuito basin (30a)” [-17.50875, 20.06594] (Conradie et al. 2016:9- 
10, 16); “Cuito basin (30d)” [-17.51327, 20.06111] (Conradie et al. 2016:9-10, 16); “Cuito basin 
(32)” [-17.04880, 19.53333] (Conradie et al. 2016:9-10, 16); “Cuito basin (35)” [-16.62322, 
19.05352] (Conradie et al. 2016:9-10, 16); “Cuando basin (43)” [-17.53500, 23.18916] (Conradie 
etal. 2016:9-10, 16); “Cubango basin (45)” [-16.88350,18.01180] (Conradie etal. 2016:9, 12, 16). 
Undetermined Locality: “areas of forest and savanna in the north and northeast of Angola” (Cei 
1977:17); “plateaus regions” (Cei 1977:17). 
Taxonomic and distributional notes: Hyperolius cinereus Monard, 1937 was described 
based on two syntype specimens from “Kalukembe” in Huila Province and “Bimbi” in Huambo 
Province (Monard 1937a). Laurent (1964a) later assigned a small collection of specimens from 
“Dundo” in Lunda Norte Province to H. cinereus, althought his characterization of the coloration 
of this species differs from that of Monard. Both Monard (1937a) and Laurent (1964a) examined 
only preserved material and could only speculate on the coloration in life (Conradie et al. 2013). 
Conradie et al. (2013) considered that populations from southern and central Angola, including the 
localities for Monard’s two syntypes, correspond to H. cinereus. However, the northern populations 
