MARQUES ET AL.; AMPHIBIANS AND TERRESTRIAL REPTILES OF ANGOLA 
lOI 
15.95000] (Bocage I866a:55, 56,1866b:76, I895a:I7I, 172, I897a:204; Laurent 1961:79; Schiotz 
1975:123, Perret I976a:25; Frost 1985:209,2016); “Kalandula” [-9.10000, 15.95000] (Conradie et 
al. 2013:206), “CangandalaNational Park” [-9.84606, 16.72233] (Ceriaco et al. 2016b:37). Kwan¬ 
za Norte: “N’golla Bumba” [-9.03333, 14.60000] (Ferreira 1906:161; Ceriaco et al. 2014a: 19); 
“Rio Luinha” [-9.26667, 14.53333] (Ferreira 1906:163); “Quilombo” [-9.33333, 14.90000] (Fer¬ 
reira 1906:163). Kwanza Sul: “Novo Redondo” [-11.20000,13.85000] (Ferreira 1906:163); “Con- 
gulu” [-10.86667, 14.28333] (Parker 1936:144). Benguela: “Hanha” [-13.30000, 14.20000] 
(Bocage 1897a:204, 1897b:211; Perret 1976a:25); “Cubal” [-13.03333, 14.25000] (Mertens 
1938:429). Undetermined Locality: “Mupepe” (Ferreira 1906:163); “areas of forest and savanna 
in the north and northeast of Angola” (Cei 1977:17); “plateaus regions” (Cei 1977:17). 
Taxonomic and distributional notes: Bocage (1866b) described two new species, Hyperolius 
cinnamomeo-ventris Bocage, 1866 and Rappia tristis (Bocage, 1866) from “Duque de Braganga [= 
Calandula].” The nomen tristis was synonymized with H. cinnamomeoventris by Laurent (1943b, 
1947) and has subsequently been followed by other authors including Perret (1976a). Before the 
Lisbon fire Perret (1976a) visited the Museu Bocage but did not find the holotype of either of these 
species. One syntype of Rappia bivittata designated by Ferreira (1906) from “N’Golla Bumba,” 
Kwanza Sul province seems to represent Hyperolius cinnamomeoventris Bocage, 1866 (Ceriaco et 
al. 2014a). Currently this species is accepted and recognized as Hyperolius cinnamomeoventris 
(Schiotz 1999; Channing 2001; Fretey et al. 2011; Conradie et al. 2013; Frost 2016). This wide¬ 
spread species represents a complex of species found in Central Africa and several offshore islands 
and is awaiting taxonomic revision. 
Hyperolius concolor (Hallowell, 1844) Variable Reed Frog 
Ixalus concolor Hallowell 1844:60. Holotype: ANSP 3216 (Malnate 1971:350) (collector unknown). Type 
locality: “Liberia, W. Africa” (Hallowell 1844:60), Liberia. 
Hyperolius modestus: Bocage (1866a:55, 1866b:74). 
Rappia concolor. Bocage (1895a:173), (1897b:211). 
Hyperolius eoneolor Noble (1924:252), Schiotz (1999:104), Amiet (2012:264), Frost (2016). 
Cameroon, and along the Atlantic coast south 
to Angola. 
Ocurrences in Angola (Map 40): The 
species is recorded mostly from western Ango¬ 
la. Malanje: “Duque de Braganga” [-9.10000, 
15.95000] (Bocage 1866a:55, 1866b:74). Bie: 
“Bihe” [-12.38333, 16.95000] (Bocage 
1895a:173). Benguela: “Hanha” [-13.30000, 
14.20000] (Bocage 1897b:211). Huila: 
“Caconda” [-13.73333, 15.06667] (Bocage 
1895a:173); “Huila” [-15.05000, 13.55000] 
(Bocage 1895a: 173). Undetermined Locality: 
“Rio Quando” (Bocage 1895a: 173). 
Taxonomic and distributional notes: 
The Angolan records of Hyperolius concolor 
(Hallowell, 1844) probably represent a 
Map 40. Distribution of Hyperolius concolor in Angola. 
Global conservation status (lUCN): Least Concern. 
Global distribution: The species is irE 
known from eastern Sierra Leone to western 
I6“E 20*E 
