106 
PROCEEDINGS OF THE CALIFORNIA ACADEMY OF SCIENCES 
Series 4, Volume 65, Supplement II 
Ocurrences in Angola (Map 46): The 
species is only known from the type locality 
“Landana” in the Cabinda enclave. Cabinda: 
“Landana” [-5.21667, 12.15000] (Rochebrune 
1885:91; Frost 1985:212, 2016). 
Taxonomic and distributional notes: 
The species was described by Rochenbrune 
(1885) from “Locum Landana dictum, in foliis 
Musarum” in the Cabinda enclave. Laurent {in 
Frost 1985), and latter Fretey et al. (2011) con¬ 
sidered Hyperolius maestus Rochebrune, 1885 
to be a possible synonym of Hyperolius 
marmoratus Rapp, 1842. The type specimen 
was deposited in Museo Bouvier and is now 
presumably in Museum national d’Histoire 
naturelle, Paris and unrecognized as a type if it 
still exists (Frost 2016). The species is not 
mentioned in Schiotz (1999) or Channing 
( 2001 ). 
Hyperolius nasutus Gunther, 1865 Large-Nosed Long Reed Frog 
Hyperolius nasutus Gunther 1865a:482, PI XXXIII, fig. 3. Holotype: BMNH 1947.2.9.68 (collector 
J.F. Bayao). Type locality: “Duque de Bragan^a” [= Calandula], Malanje Province, Angola. 
Rappia punctulata Bocage 1895a: 168. Holotype: MBL T.23-227 (collector Banyures) destroyed by fire 18 
March 1978. Type locality: “sur les bords du Quanza” [= Quanza river], Angola. 
Hyperolius nasutus'. Bocage (1866a:55), Noble (1924:253), Schmidt (1936:132), Loveridge (1936a: 109, 
1953a:362, 1953:362, 1957:333), Monard (1937a:39, 1938:94), Mertens (1938:429), Frade (1963:254), 
Haacke (1970:279), Schiotz (1975:97), Poynton and Broadley (1976:206), Cei (1977:17, 18), Frost 
(1985:214, 2016), Poynton and Haacke (1993:14), Schiotz (1999:97, 2006:65), Channing (2001:166), 
Largen (2001:359), Channing et al. (2002:96), Schiotz and Van Daele (2003:138), Amiet (2005:275f 
Conradie et al. (2012a:3), Ceriaco et al. (2016b:35), Conradie et al. (2016:11). 
Rappia nasuta: Boulenger (1882:127), Bocage (1895a: 169, 1897a:204). 
Rappiapunetulata: Bocage (1897a:204), Ferreira (1904:112), Amiet (2005:275). 
Hyperolius punetulatus: Noble (1924:253), Loveridge (1936a: 107, 1936b:405), Mertens (1937:20), Monard 
(1938:86), Frade (1963:254), Perret (1976a:27), Frost (1985:216). 
Hyperolius nasutus nasutus'. Laurent (1950a: 17, 1954a:84, 1964a: 154), Hellmich (1957a:29). 
Hyperolius {Hyperolius) nasutus'. Fretey et al. (2011:32). 
Global conservation status (lUCN): Least Concern. 
Global distribution: The species is known from Angola south to the Okavango Delta of 
Botswana and adjacent northern Namibia. 
Ocurrences in Angola (Map 47): The species is known from its type locality “Duque de Bra- 
ganqa, Malanje Province” and is very widespread across almost the whole of the country. Bengo: 
“Cabiri” [-8.91667, 13.66667] (Ferreira 1904:112). Kwanza Norte: “3 km W of Salazar” 
[-9.30000, 14.91667] (Poynton and Haacke 1993:14); “Dondo” [-9.68333, 14.43333] (Loveridge 
1936a:107). Kwanza Sul: “Namba” [-11.91667, 14.85000] (Poynton and Haacke 1993:14). 
Malanje: “Duque de Braganqa” [-9.10000, 15.95000] (Gunther 1865a:482; Bocage 1866a:55, 
1895a:169, 1897a:204; Boulenger 1882:127; Loveridge 1936a:109, 1936b:405, 1953a:362, 
1957:333; Schiotz 1975:97; Frost 1985:214, 2016; Amiet 2005:275); “Calandula” [-9.07917, 
