MARQUES ET AL.: AMPHIBIANS AND TERRESTRIAL REPTILES OF ANGOLA 
107 
15.79583] (Channing et al. 2013:334); “Kan- 
gandala” [-9.825II, 16.91225] (Channing etal. 
2013:334); “Cangandala National Park” 
[-9.84606,16.72233] (Ceriacoetal. 2016b:35); 
Lunda Norte: “Dundo” [-7.36667, 20.83333] 
(Laurent 1954a:84); “environs de Dundo, rives 
de la Luaehimo” [-7.38333, 20.85000] (Lau¬ 
rent 1950a: 17); “Carumbo lagoon” [-7.74422, 
19.95467] (Conradie et al. 2012a:3); “Muita 
(Luembe E)” [-7.80000, 21.45000] (Laurent 
1950a: 17); “Kossa a 100 km au sud-est de 
Dundo” [-7.90000, 21.36667] (Laurent 
1950a: 17); “rive de la Tehihumbwe 40 km a 
Test de Dundo” [-8.01667, 19.31667] (Laurent 
1950a:17); “Village Capaia” [-8.33847, 
20.24250] (Conradie et al. 2012a:3). Lunda 
Sul: “Dala” [-11.03333, 20.20000] (Monard 
1937a:39, 1938:94); “Alto Chieapa,sourees du 
Cuilo” [-10.88333, 19.23333] (Laurent 1964a:154); “Alto Cuilo, rives du Cuilo “ [-10.01667, 
19.55000] (Laurent 1964a:154). Mexico: “region du lae Calundo” [-11.80000, 20.86667] (Laurent 
1964a:154); “reserve de ehasse de Cameia” [-11.83333,21.00000] (Laurent 1964a:154). Bie: “Chi- 
tau” [-11.43333, 17.15000] (Sehmidt 1936:132); “General Maehado” [-12.01667, 17.06667] Mer- 
tens (1937:20). Huambo: “Bimbi” [-11.81667, 15.83333] (Monard 1937a:39, 1938:94); “Tongru- 
be am Jamba Fliipehen” [-13.60000, 16.60000] (Hellmieh 1957a:29). Benguela: “Cubal” 
[-13.03333,14.25000] (Mertens 1938a:429); “Mareo de Canavezes (Cubal da Ganda)” [-13.08333, 
14.33333] (Laurent 1964a:154). Hufla: “Huilla” [-15.05000, 13.55000] (Boeage 1895a:169, 
1897a:204); “Caeonda” [-13.73333, 15.06667] (Boeage 1895a:169, 1897a:204); “Kalukembe” 
[-15.05000, 13.55000] (Monard 1938:86); “Nunteehite lagoon” [-15.13333, 13.41667] (Poynton 
andHaaeke 1993:14). Cunene: “Chimporo” [-16.03333, 17.15000] (Monard 1937a:39, 1938:94). 
Cuando Cubango: “Kuandu” [-16.74487, 19.10136] (Monard 1937a:39, 1938:94). Undeter¬ 
mined Locality: “Margens do Cuanza” (Boeage 1895a: 168, 1897a:204; Loveridge 1936a: 107, 
1957:333; Perret 1976a:27; Frost 1985:216; Amiet 2005:275); “without preeise loeation” (Monard 
1937a:39, 1938:94); “Western subregion and Angolan highlands (Angolan eoastal or watersheds)” 
(Frade 1963:254); “Zambezian highlands” (Frade 1963:254); “areas of forest and savanna in the 
north and northeast of Angola” (Cei 1977:17); “plateaus regions” (Cei 1977:17); “arid territories 
along the eoast” (Cei 1977:18). 
Taxonomic and distributional notes: While the type speeimen of R. punctulata is now lost, 
Channing et al. (2013) ineluded it as a synonym of H. nasutus, for whieh there are many junior 
synonyms. To date, fifteen names have been used for members of the nasutus group, most of whieh 
have been synonymized with the nominotypieal form (Sehiotz 1999); a relatively reeent list of 
these synonyms is provided by Amiet (2005). Poynton and Broadley (1987) reeognized three 
speeies in the southern Afriean savanna, Hyperolius viridis Sehiotz, 1975, Hyperolius nasutus 
Gunther, 1865 and Hyperolius benguellensis Boeage, 1893. Sehiotz (1999) ineluding//, benguel- 
lensis as a synonym of H. nasutus after arguing that differenees in morphology and eolor pattern 
are not suffieient for distinguishing these two speeies. Later, Channing et al. (2002) retained 
H. benguellensis as a synonym of nasutus, but also divided the group into three eryptie speeies 
differing in advertisement eall: Hyperolius acuticeps Ahl, 1931, Hyperolius viridis Sehiotz, 1975, 
1,2°E 16°E 20°E H°E 
Map 47. Distribution of Hyperolius nasutus in Angola. 
