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PROCEEDINGS OF THE CALIFORNIA ACADEMY OF SCIENCES 
Series 4, Volume 65, Supplement II 
and H. nasutus (Schiotz and Van Daele 2003; Schiotz 2006). Schiotz (2006) distinguished two 
speeies, H. benguellensis and H. nasutus, in the nasutus group for Angola based on the strueture of 
the advertisement call and color pattern. This widespread species is common in savannas and 
exhibits a preference for humid areas such as reeds growing in deep water, pools, streams, and large 
rivers (Schiotz 1999; Channing 2001). 
HyperoUus nitidulus Peters, 1875 Plain Reed Frog 
Hyperolius nitidulus Peters 1875:209, pi. 3, figs. 4, 4a. Holotype: ZMB 7729 fide Bauer et al. (1995:44) (eol- 
leetor R. Buehholz). Type loeality: “Yomba (Lagos)” (Peters 1875:209), Nigeria. 
Hyperolius nitidulus: Peters (1877a:619), Sehiotz (1999:206), Wieezorek et al. (2000:1238-1241), Rodel et al. 
(2010:178), Amiet (2012:194), Frost (2016). 
Global conservation status (lUCN): 
Least Concern. 
Global distribution: The species is 
known from West African extending from 
Guinea east into Nigeria, Angola and 
Cameroon. 
Ocurrences in Angola (Map 48): The 
species was cited from “Chinchoxo” in the 
Cabinda enclave. Cabinda: “Chinchoxo” 
[-5.10000, 12.10000] (Peters 1877a:619). 
Taxonomic and distributional notes: 
According to Wieezorek et al. (2000), the 
species Hyperolius nitidulus Peters, 1875 is a 
West African species associated with savanna 
and woodlands (Schiotz 1999; Wieezorek et al. 
2000; Rodel et al. 2010; Amiet 2012). Based on 
the distribution of the species delimited by 
Wieezorek et al. (2000), the record from north¬ 
western Angola is likely a misidentification. 
Hyperolius ocellatus (Gunther, 1858) 
Hyperolius oeellatus Gunther 1858a:326. Lectotype: BMNH 1947.2.9.22, (collector L. Fraser), formerly 
BMNH 51.11.13.48, by implication of being referred to as “holotype” by Perret (1975a:27), Syntypes: 
BMNH [2 specimens] (collector Mr. Fraser, for the “Fernando Po” specimen) (Gunther 1858a:326). Type 
locality: “Fernando Po” [= Bioko, Guinea] fide Perret (1975a:24), who considered “Angola” an error. 
Originally given as “Fernando Po” and “Angola” by Gunther (1858a:326). 
Hyperolius Lueani Rochebmne 1885:91. Type: Presumably in MNHP (collector unknown), uncatalogued as 
a type, if still extant, formerly in Museo Bouvieri. Type locality: “Locam Landana dictum, in frondibus 
Borassomm” [= Landana, Cabinda enclave] Cabinda enclave, Angola. 
Hyperolius oeellatus: Gunther CT858b” 1859:88), Perret (1975a:24), Cei (1977:17), Frost (1985:214, 2016), 
Schiotz (1999:141), Channing (2001:168), Channing et al. (2012:217). 
Rappia oeellata: Boulenger (1882:123); Bocage (1895a: 165). 
Hyperolius lueani: Frost (1985:212, 2016). 
Hyperolius {Hyperolius) oeellatus: Fretey et al. (2011:32). 
Global conservation status (lUCN): Least Concern. 
Global distribution: The species is from tropical Central Africa. It occurs in the Cabinda 
irE I6®E 20°E :4»E 
Map 48. Distribution of Hyperolius nitidulus in Angola. 
Golden-Eyed Reed Frog 
