112 
PROCEEDINGS OF THE CALIFORNIA ACADEMY OF SCIENCES 
Series 4, Volume 65, Supplement II 
ned Locality: “bords du Quanza” (Bocage 1895a: 173, 1897:204); “Rio Quando” (Bocage 
1895a:173, 1897a:204); “Zambezian highlands” (Frade 1963:254); “plateaus regions” (Cei 
1977:17). 
Taxonomic and distributional notes: Boulenger (1882) noted that this is an East African 
species, although he recognized (Bocage 1866b) Hyperolius microps as a valid name. Based on this 
information, the Angolan records possibly belong to another Hyperolius species and it would be 
important to review this case. Unfortunately, the specimens from Museu Bocage (Bocage 1866b, 
1895a, 1897a) were destroyed in the 1978 fire, although the specimens studied by Boulenger 
(1905) still exist in the British Museum. Noble (1924) also cited specimens of H. pusillus and 
H. microps in the collections of the American Museum of Natural History from Angola, though 
without precise locality information. 
Hyperolius quinquevittatus Bocage, 1866 Five-Striped Reed Frog 
Hyperolius quinquevittatus Bocage 1866a:56. Syntypes: MBL 24-214, 24-215 [2 specimens] (collector F.A.P. 
Bayao). Type locality: “Duque de Braganga” [= Calandula], Malanje Province, Angola. 
Hyperolius quinquevittatus: Bocage (1866h:77), Frade (1963:254), Noble (1924:253), Laurent (1950A:16, 
1954a:79), Perret (1976a:25), Cei (1977:17), Frost (1985:216, 2016), Schiotz (1999:101), Channing 
(2001:178), Channing and Howell (2006:180). 
Rappia quinquevittata: Bocage (1895a:174). 
Hyperolius quinquevittatus quinquevittatus: Schiotz (1975:102), Poynton and Broadley (1987:197), Pickers- 
gill (2007a:360). 
Hyperolius {Hyperolius) quinquevittatus: Fretey et al. (2011:33). 
According to Perret (1976a), the syntypes 
described by Bocage (1866a), which were 
destroyed in the 1978 fire in Lisbon, were stud¬ 
ied by Schiotz (1975) when he concluded that 
H. quinquevittatus is similar in morphology to 
H. nasutus Gunther, 1865. Poynton and 
Broadley (1987) noted, however, that Hyper¬ 
olius quinquevittatus is more similar to Hyper¬ 
olius kivuensis even if H. quinquevittatus is in 
general much smaller and slimmer. Pickersgill 
Global conservation status (lUCN): Least Concern. 
Global distribution: The species is known from open savannas at higher elevations in north¬ 
ern Angola, the southern Democratic Republic of Congo, southern Tanzania, and northern Zambia 
and Malawi, and Mt. Namuli in northern Mozambique. 
Ocurrences in Angola (Map 54): The species occurs especially in northern Angola. Malan¬ 
je: “Duque de Braganqa” [-9.10000, 15.95000] 
(Bocage 1866a:56, 1866b:77, 1895a:174; 
Schiotz 1975:102; Perret 1976a:25; Frost 
1985:216, 2016; Poynton and Broadley 
1987:197; Pickersgill 2007a:360). Undeter¬ 
mined Locality: “areas of forest and savanna 
in the north and northeast of Angola” (Cei 
1977:17); “plateaus regions” (Cei 1977:17). 
Taxonomic and distributional notes: 
Map 54. Distribution of Hyperolius quinquevittatus in 
Angola. 
