MARQUES ET AL.; AMPHIBIANS AND TERRESTRIAL REPTILES OF ANGOLA 
II3 
(2007a) suspected that the species belongs to the multifasciatus group, while Fretey et al. (2011) 
included the H. multifasciatus in the synonym of quinquevittatus. 
Hyperolius raymondi Conradie, Branch and Tolley, 2013 Raymond’s Reed Frog (Endemic) 
Hyperolius raymondi Conradie et al. 2013:203, fig. 7. Holotype: PEM A10049 (eollectors W. Conradie, 
W. Branch, P. Vaz Pinto, S. Batista and N. Batista). Type locality: “Lagoa Cammbo from a dambo near the 
expedition base camp,” Lunda Norte Province, Angola. 
Hyperolius cinereus: Laurent (1964a: 149), Schiotz (1999:192), Channing (2001:152). 
Hyperolius {Hyperolius) einereus: Fretey et al. (2011:30). 
Hyperolius raymondi'. Frost (2016). 
Global conservation status (lUCN): Data Deficient. 
Global distribution: The species is known from Angola. 
Ocurrences in Angola (Map 55): The 
species is known from the type locality “Lagoa 
Cammbo” and from surrounding localities in 
Lunda Norte Province. Lunda Norte: 
“Dundo” [-7.36667, 20.83333] (Laurent 
1964a: 149; Channing 2001:153; Conradie et 
al. 2013:203); “margins Lagoa Samokaza east 
of Lagoa Cammbo” [-7.73833, 19.98694] 
(Conradie et al. 2013:203); “Lagoa Cammbo 
from a dambo near the expedition base camp” 
[-7.74422, 19.95522] (Conradie et al. 
2013:203; Frost 2016); “headwaters of Lovua, 
north of village Capaia” [-8.33847, 20.24250] 
(Conradie et al. 2013:203). 
Taxonomic and distributional notes: 
Populations that Laurent (1964a) referred to 
H. cinereus were recognized by Conradie et al. 
(2013a) as a distinct species and described as 
Map 55. Distribution of Hyperolius raymondi in Angola. 
H. raymondi (see further discussion above in account for H. cinereus). This newly described 
species is so far endemic to Angola, but may also occur in southernmost Democratic Republic of 
Congo and northwest Zambia. It is found in grass-covered “dambo” (water levels less than 30 cm) 
in the flood plains of the Lulele River, a southern tributary draining north to the larger Congo River 
(Conradie et al. 2013). 
Hyperolius rhizophilus Rochebrune, 1885 African Reed Frog (Endemic) 
Hyperolius Rhizophilus Rochebmne 1885:92. Type: Presumably in MNHN (eolleetor unknown), uneata- 
logued as a type, if still extant, formerly in Museo Bouvieri. Not eited by Guibe (1950). Type locality: 
“Locum Landana dictum, in foliis rhizophoramm” [= Landana, Cabinda enclave], Cabinda Province, 
Angola. 
Hyperolius rhizophilus Rochebmne, 1885: Frost (1985:215, 2016). 
Hyperolius {Hyperolius) rhizophilus Rochebmne, 1885: Fretey et al. (2011:33). 
Global conservation status (lUCN): Data Deficient. 
Global distribution: The species is known only from Angola. 
Ocurrences in Angola (Map 56): The species is only known from the type locality “Landana” 
