MARQUES ET AL.; AMPHIBIANS AND TERRESTRIAL REPTILES OF ANGOLA 
II9 
1964a: 145); “Alto Cullo, Poste de Cacolo, galerie forestiere du Tcha-Muchito, sous-affluent du 
Cullo” [-10.01667, 19.45000] (Laurent 1964a;145); “Alto Chicapa, galerie forestiere de laNgun- 
go, sous-affluent du Cuango-Muque (Rio Gungo)” [-10.83333, 19.28333] (Laurent 1964a: 145; 
Ruas 1996:28); “Alto Chieapa, Tshimboma, affl. rive gauehe du Cuango-Muque” [-10.76667, 
19.20000] (Laurent 1964a: 145; Ruas 1996:28); “Alto Cuilo, rives du Cuilo (Rio Cuilo)” 
[-10.86667, 19.40000] (Laurent 1964a: 145; Ruas 1996:28); “Alto Chieapa, Caseade de la Kamu- 
tolonga (Rio Camuntongola)” [-10.88333, 19.25000] (Laurent 1964a: 145; Ruas 1996:28); “Alto 
Chieapa, sourees de la Tshimboma, affl. rive gauehe du Cuango-Muque (Rio Chimboma)” 
[-10.88333, 19.23333] (Laurent 1964a: 146; Ruas 1996:28). Undetermined Locality: without 
precise locality (Laurent 1964a: 146), “areas of forest and savanna in the north and northeast of 
Angola (Cei 1977:16). 
Taxonomic and distributional notes: Channing (2001) provided two unspecified localities 
for A. lameerei in northwestern Angola, and he noted that when collected in Angola this species 
was found in in leaf litter. 
Arthroleptis spinalis Boulenger, 1919 Tanganyika Screeching Frog 
Arthroleptis spinalis Boulenger 1919b: 187. Holotype: MRAC (eolleetor Stappers), fide R. Laurent in Frost 
(1985:20). Type loeality: “laplaine Saint-Louis, auLake Tanganyika” (Boulenger 1919b: 187), [= St. Louis 
Plain on the western shore of Lake Tanganyika] Demoeratie Republie of Congo. 
Athroleptis boulengeri: Laurent (1950a: 15). 
Schoutedenella spinalis: Cei (1977:16). 
Arthroleptis spinalis: Fretey et al. (2011:22), Frost (2016). 
Global conservation status (lUCN): Data Deficient. 
Global distribution: The species is known from northern Angola and the eastern Democratic 
Republic of Congo. 
Ocurrences in Angola (Map 64): The 
species has one record in the northeast of the 
country. Lunda Norte: “Dundo, galerie fores¬ 
tiere de la Luachimo” [-7.38333, 20.83333] 
(Laurent 1950a: 15). 
Taxonomic and distributional notes: 
Boulenger (1919a) described Arthroleptis 
spinalis from eastern Democratic Republic of 
Congo, which is generally recognized as the 
only known locality for this species (Fretey et 
al. 2011, Frost 2016). However, Laurent 
(1950a) cited Arthroleptis boulengeri (de 
Witte, 1921) from northeastern Angola and 
then later recognized this species as a junior 
synonym of A. spinalis (Laurent 1954). If cor¬ 
rect, this suggests that this species may be more 
widespread throughout northern Angola and 
southern Democratic Republic of Congo. ^4. Distribution of Arthroleptis spinalis in Angola. 
12*E l(i*E 2I)°E 24*E 
