120 
PROCEEDINGS OF THE CALIFORNIA ACADEMY OF SCIENCES 
Series 4, Volume 65, Supplement II 
Arthroleptis stenodactylus Pfeffer, 1893 Common Squeaker 
Arthroleptis stenodactylus Pfeffer 1893:93, pi. 1, fig. 11. Holotype: ZMH, presumably destroyed in World War 
II (eolleetor F. Stuhlmann). Type loeality: “Kihengo” (Pfeffer 1893: pi. I), Tanzania. 
Corachodichus stenodactylus: Frade (1963:254), Cei (1977:16). 
Corachodichus stenodactylus stenodactylus: Laurent (1964a: 144). 
Arthroleptis stenodactylus: Ruas (1996:28, 2002:145), Channing (2001:46), Fretey et al. (2011:25), Frost 
(2016). 
Global conservation status (lUCN): Least Concern. 
Global distribution: The species is broadly distributed from Kenya and Tanzania, west into 
the Democratic Republic of Congo and Angola, and south through Zambia, Zimbabwe, Malawi, 
Mozambique, and extreme northeastern South Africa. 
Ocurrences in Angola (Map 65): The 
species is known only for Moxico Province. 
Moxico: “Calombe, Luso” [-11.83333, 
19.93333] (Ruas 1996:28, 2002:145; Channing 
2001:47); “Cazombo” [-11.88333, 22.91667] 
(Laurent 1964a; 144; Ruas 1996:28; Channing 
2001:47); “Calunda, Alto Zambeze” 
[-12.11667, 23.46667] (Laurent 1964a:144; 
Ruas 1996:28; Channing 2001:47). Undeter¬ 
mined Locality: “areas of forest and savanna 
in the north and northeast of Angola” (Cei 
1977:16); “Zambezian highlands” (Frade 
1963:254). 
Taxonomic and distributional notes: 
According to Ruas (1996), Arthroleptis sten¬ 
odactylus Pfeffer, 1893 in Angola occurs only 
in the east of the country. Previous authors 
(e.g., Channing and Howell 2006; Pickersgill 
2007a) have suggested that multiple distinct 
species exist within what is referred to A. stenodactylus, and preliminary molecular genetic data 
provides some support for this (Blackburn 2008). It is not yet clear to which of these cryptic species 
these Angolan populations might correspond. 
Map 65. Distribution of Arthroleptis stenodactylus in 
Angola. 
Arthroleptis xenochirus Boulenger, 1905 Plain Squeaker 
Arthroleptis xenochirus Boulenger 1905:108, pi. 4, figs. 2, 2a. Holotype: BMNH 1947.2.30.54 (eolleetor 
W.J. Ansorge), formerly 1904.5.2.101, R. Laurent in Frost (1985:20). Type loeality: “Marimba” 
(Boulenger 1905:109), Angola. 
Schoutedenella xenochira: Laurent (1954e:36). 
Schoutedenella xenochirus: Cei (1977:16), Laurent (1964a:145), Frost (1985:20), Channing (2001:52). 
Arthroleptis xenochirus: Laurent (1950a: 15), Frade (1963:254), Ruas (1996:28, 2002:145), Fretey et al. 
(2011:25), Frost (2016), Ceriaeo et al. (2016b:23). 
Global conservation status (lUCN): Least Concern. 
Global distribution: A wide-ranging species found in northern Angola, the southern Demo¬ 
cratic Republic of Congo, northern Zambia and possibly southwestemmost Tanzania. 
Ocurrences in Angola (Map 66): The species occurs in northeastern Angola. Lunda Norte: 
“Dundo” [-7.36667,20.83333] (Laurent 1950:15). Lunda Sul: “Poste de Cacolo, Alto Cuilo, gale- 
