MARQUES ET AL.; AMPHIBIANS AND TERRESTRIAL REPTILES OF ANGOLA 
I3I 
em regions of the country. Lunda Norte: 
“Muita (Luembe E)” [-7.80000, 21.45000] 
(Laurent 1950a: 14; Ruas 1996:26). Lunda 
Sul: “Dala” [-11.03333, 20.20000] (Monard 
1937a:52, 1938:110; Ruas 1996:26). Kwanza 
Sul: “Congulu” [-10.86667, 14.28333] (Parker 
1936:142; Ruas 1996:26). Huambo: “Bimbi” 
[-11.81667, 15.83333] (Monard 1937a:52, 
1938:110; Ruas 1996:26); “Mt. Moco” 
[-12.41667, 15.18333] (Parker 1936:14; Ruas 
1996:26). Benguela: “Benguela” [-12.58333, 
13.41667] (Loveridge 1936a:95). HuiTa: 
“Kalukembe” [-13.78333, 14.68333] (Monard 
1937a:52, 1938:110; Ruas 1996:26). Cuando 
Cubango: “Kandingu (Kului)” [-14.66667, 
17.70000] (Monard 1937a:52, 1938:110; Ruas 
1996:26). Undetermined Locality: “Between 
Benguella and Bihe” (Boulenger 1905:107; 
Map 79. Distribution of Ptychadena ansorgii in Angola. 
Loveridge 1933:371, 1936b:419, 1953a:373, 1957:343; Schmidt and Inger 1959:61; Poynton and 
Broadley 1985b: 152; Frost 1985:471, 2016; Ruas 1996:26), “areas of forest and savanna in the 
north and northeast of Angola” (Cei 1977:16); “plateaus regions” (Cei 1977:17). 
Taxonomic and distributional notes: Relatively little is known of the biology of this species, 
which is found in the forests and savannas of south-central Africa. 
Ptychadena bunoderma (Boulenger, 1907) Rough Grass Frog 
Rana bunoderma Boulenger 1907a:214. Holotype: BMNH 1947.2.2.86 (eolleetor W.J. Ansorge). Type loeal- 
ity: “Caeonda,” Huila Provinee, Angola. 
Rana (Ptychadena) buneli: Monard (1937a:55, 1938: 
A. Monard). Type loeality: “environs de Dala, 
sur le Tyiumbwe,” Lunda Sul Provinee, Angola. 
Rana bunoderma'. Sehmidt (1936:129). 
Ptychadena bunoderma'. Laurent (1964a: 142), Cei 
(1977:16), Poynton and Broadley (1985b: 152), 
Frost (1985:471, 2016), Ruas (1996:26), Largen 
(2001:177), Channing (2001:324). 
Ptychadena (Ptychadena) bunoderma'. Fretey et al. 
(2011:41). 
Global conservation status (lUCN): 
Least Concern. 
Global distribution: The species is 
known from Angola and northwestern Zambia. 
Ocurrences in Angola (Map 80): The 
species occurs in central and eastern Angola. 
Lunda Sul: “Alto Chicapa” [-10.88333, 
19.23333] (Laurent 1964a: 142; Ruas 1996:26); 
“Alto Chicapa, humidiherbosa des sources de 
la Kamassaka (Rio Camassaca)” [-10.88333, 
114). Syntypes: MNHNC [three speeimens] (eolleetor 
12°E l(i®E 20°E I4'*E 
Map 80 Distribution of Ptychadena bunoderma in 
Angola. 
