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PROCEEDINGS OF THE CALIFORNIA ACADEMY OF SCIENCES 
Series 4, Volume 65, Supplement II 
19.25000] (Laurent 1964a: 142; Ruas 1996:26); “Alto Chicapa, humidiherbosa des sources de la 
Kamutongola (Rio Camuntongola)” [-10.53000, 19.15000] (Laurent 1964a: 142; Ruas 1996:26); 
“environs de Dala, sur le Tyiumbwe (Rio Chiumbe)” [-11.03333, 20.20000] (Monard 1937a:55, 
1938:114; Ruas 1996:26). Bie: “Chitau” [-11.43333, 17.15000] (Schmidt 1936:129; Ruas 
1996:26). Hmla: “Caconda” [-13.73333, 15.06667] (Boulenger 1907a:214; Poynton and Broadley 
1985b: 152; Frost 1985:471, 2016; Ruas 1996:26). Undetermined Locality: “areas of forest and 
savanna in the north and northeast of Angola” (Cei 1977:16). 
Taxonomic and distributional notes: Gaston de Witte initially identified Monard’s types of 
R. buneli as ''Rana n. sp., voisine de Rana mascareniensis D.B.,” although based on Monard’s 
(1937a) description it is closer to P. bibronii or P. bunoderma. Laurent (1964a) provided some new 
records for Ptychadena bunoderma in Angola, including from near the type locality of 
P. buneli which he placed in the synonymy of P. bunoderma. Channing (2001) provided several 
records further north for the species without detailed information. 
Ptychadena grandisonae Laurent, 1954 Grandison’s Grass Frog 
Ptychadena grandisonae Laurent 1954b: 11, pi. 1, figs. 2, 5, pi. 3, figs. 1-2, pi. 4, figs. 1, 9. Holotype: pre¬ 
sumably MD 506? (eolleetor M. de Petehkowsky). Type loeality: “Muita, Luembe E” [= Luembe] Lunda 
Norte, Angola. 
Rana (Ptyehadena) bibroni: Monard (1937a:51, 1938:109). 
Ptyehadena bibronii'. Laurent (1950a: 14). 
Ptyehadena grandisonae'. Laurent (1964a: 139), Cei (1977:16), Poynton and Broadley (1985b: 150), Frost 
(1985:472, 2016), Poynton and Haaeke (1993:14), Channing (2001:325). Ruas (2002:144). 
Rana grandisonae'. Sehmidt and Inger (1959:70). 
Ptyehadena (Ptyehadena) grandisonae'. Fretey et al. (2011:41). 
Ptyehadena ef grandisonae'. Conradie et al. (2016:17). 
Global conservation status (lUCN): Least Concern. 
Global distribution: The species is known from the Democratic Republic of Congo, Angola, 
and Zambia. 
Ocurrences in Angola (Map 61): The 
species occurs in the northeast of the country, 
although there are some records further south. 
Lunda Norte: “environs du Dundo” 
[-7.36667, 20.83333] (Laurent 1964a: 139); 
“Muita (Luembe E)” [-7.80000, 21.45000] 
(Laurent 1950a: 14, 1954b: 11; Schmidt and 
Inger 1959:70; Poynton and Broadley 
1985b: 150; Frost 1985:472,2016). Lunda Sul: 
“Alto Chicapa, mare Tcha-Mutuka, pres des 
sources du Cuango-Muque” [-10.76667, 
19.20000] (Laurent 1964a:139); “Alto Chica¬ 
pa, sources du Cuilo” [-10.86667, 19.40000] 
(Laurent 1964a: 139); “sources de la Kamuton¬ 
gola, Alto Chicapa” [-10.53000, 19.15000] 
(Laurent 1964a:139); “Lunda” [-10.96667, 
20.06667] (Monard 1937a:51, 1938:109). 
Mexico: “Luso” [-11.78333, 19.91667] (Ruas 
2002:144). Benguela: “Ebanga” [-12.73333, 
12°E l(j®E 2()°E I4"E 
Map 81. Distribution of Ptychadena grandisonae in 
Angola. 
