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PROCEEDINGS OF THE CALIFORNIA ACADEMY OF SCIENCES 
Series 4, Volume 65, Supplement II 
Taxonomic and distributional notes: Poynton and Broadley (1985b) elevated P. guibei to a 
full species. Fretey et al. (2011:41) cited Ptychadena (Ptychadena) chrysogaster Laurent, 1954 for 
Angola though we believe that this should be referred to P. guibei (Frost 2016). New material cited 
by Conradie et al. (2016) extends the species distribution into extreme southeastern Angola. 
Ptychadena keilingi Monard, 1937 Keiling’s Grass Frog 
Rana {Ptychadena) keilingi Monard 1937a:53, figs. 14-16. Syntypes: MNHC 90.0829 and 90.0830 (collector 
A. Monard). Type locality: “Dala,” Lunda Sul, Angola. 
Rana (Ptyehadena) keilingi'. Monard (1938:112), Marx (1959:442), Laurent (1964a: 141). 
Ptyehadena keilingi'. Forcart (1946:126), Cei (1977:16), Poynton and Broadley (1985b: 154), Frost (1985:472, 
2016), Ruas (1996:26), Largen (2001:177), Channing (2001:327). 
Ptyehadena {Ptyehadena) keilingi'. Fretey et al. (2011:41). 
Global conservation status (lUCN): Least Concern. 
Global distribution: The species is known from the Democratic Republic of Congo and adja¬ 
cent northeastern Angola to extreme northwestern Zambia. 
Ocurrences in Angola (Map 83): The species occurs in northeastern Angola. Lunda Norte: 
“Riviere Luita river, Poste de Cuilo (Rio 
Luita)” [-8.03333, 19.41667] (Laurent 
1964a: 141; Ruas 1996:27); “Poste de Luangue, 
humidiherbosa du risseau Katcheleka, affl. 
ouest Luangue, entre la Lunguena et le Tcha- 
Pemba (Posto do Luangue)” [-9.08333, 
19.71667] (Laurent 1964a:141; Ruas 1996:27). 
Lunda Sul: “Alto Chicapa” [-10.88333, 
19.23333] (Laurent 1964a:141; Ruas 1996:27); 
“Dala” [-11.03333, 20.20000] (Monard 
1937a:53, 1938:112; Forcart 1946:126; Marx 
1959:442; Poynton and Broadley 1895b: 154; 
Frost 1985:472,2016; Ruas 1996:26). Moxico: 
“rives du lac Calundo (Lago Calundo)” 
[-11.80000, 20.86667] (Laurent 1964a:141; 
Ruas 1996:27). Undetermined Locality: 
“areas of forest and savanna in the north and 
northeast of Angola” (Cei 1977:16). Distribution of Ptychadena keilingi in Angola. 
Taxonomic and distributional notes: Two of the five type specimens noted by Monard 
(1937a) remain in the Musee d’Histoire Naturelle, La-Chaux-de Fond, Switzerland. Little is known 
of this frog species. 
Ptychadena cf. mascareniensis (Dumeril and Bibron, 1841) Mascarene Grass Frog 
Rana maseareniensis Dumeril and Bibron 1841:350. Syntypes: MNHN 4379-4381 fide Guibe (1950), MCZ 
A-1044 (exchange from MNHN) fiide Barbour and Loveridge (1929), USNM 10975 (2 specimens) fiide 
Cochran (1961) (collectors J.R.C. Quoy and J.P. Gaimard, J.-J. Dussumier, J.-B. Leschenault, Nivoy). 
Type locality: “lies Mascareignes, ou les Sechelles, Maurice et Bourbon.” Restricted to “Island of Reu¬ 
nion, Mascarene Islands (Indian Ocean)” by Cochran (1961) and “He Bourbon” [= Reunion] by Blom- 
mers-Schlosser and Blanc (1991). 
Ptyehadena maseareniensis hylaea Schmidt and Inger 1959:83). Holotype: FMNH 57965 (collector H.A. 
Beatty). Type locality: “Mount Nimba, Liberia” Liberia. 
