MARQUES ET AL.; AMPHIBIANS AND TERRESTRIAL REPTILES OF ANGOLA 
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Rana mascareniensis: Boulenger (1905:107). 
Rana (Ptychadena) mascareniensis'. Monard (1937a:50, 1938:108). 
Rana maseareniensis maseareniensis: Loveridge (1957:342), Ruas (1996:25, 2002:144). 
Ptyehadena maseareniensis'. Cei (1977:16, 17), Frost (1985:473), Charming (2001 329), Pickersgill 
(2007a: 128), Frost (2016), Conradie et al. (2016:18). 
Ptyehadena maseareniensis bibroni: Perret (1979:6). 
Ptyehadena hylaea: Lamotte (1967:647), Pickersgill (2007a: 134). 
Ptyehadena (Ptyehadena) maseareniensis'. Fretey et al. (2011:41). 
Global conservation status (lUCN): Least Concern. 
Global distribution: Populations from throughout sub-Saharan Africa currently referred to 
this species likely represent multiple undescribed species or should be referred to other described 
species in this genus. 
Ocurrences in Angola (Map 84): The 
species is widespread across the country. 
Lunda Sul: “Dala” [-11.03333, 20.20000] 
(Monard 1937a:50, 1938:108, Ruas 1996:25). 
Moxico: “Rio Caluando” [-11.46667, 
17.70000] (Ruas 1996:25, 2002:144); “Lago 
Dilolo” [-11.50000, 22.01667] (Ruas 1996:25, 
2002:144); “Lago Cameia” [-11.71667, 
20.80000] (Ruas 1996:25, 2002:144). Bie: 
“Chitau” [-11.43333, 17.15000] (Schmidt and 
Inger 1959:84; Ruas 1996:25). Huila: “Huila” 
[-11.71667, 20.80000] (Schmidt and Inger 
1959:84). Cunene: “Chimporo” [-16.03333, 
17.15000] (Monard 1937a:50, 1938:108; Ruas 
1996:25). Cuando Cubango: “Cuabango 
basin (29)” [-17.87291,19.83333] (Conradie et 
al. 2016:9-10,18); “Cuando basin (39)” 
[-17.46333, 22.86638] (Conradie et al. 2016:9- . ^4. Distribution of Ptyehadena cf. mascareniensis 
10,18); “Cubango basin (41a)” [-17.46777, 
23.06667] (Conradie et al. 2016:9-10,18); “Cubango basin (43)” [-17.53500, 23.18916] (Conradie 
et al. 2016:9-10,18); “Cubango basin (44a)” [-17.57333, 23.26000] (Conradie et al. 2016:9-10,18); 
“Cuito basin (63)” [-17.93611, 21.10269] (Conradie et al. 2016:9,12,18). Undetermined Locali¬ 
ty: “Between Benguella and Bihe (Entre Benguela e Bie)” (Boulenger 1905:107; Ruas 1996:25); 
“Angola Quest?” (Perret 1979:9); “areas of forest and savanna in the north and northeast of Ango¬ 
la” (Cei 1977:16); “arid territories along the coast” (Cei 1977:17); “plateaus regions” (Cei 
1977:17). 
Taxonomic and distributional notes: Ptychadena mascareniensis is a complex of multiple 
cryptic forms, possibly representing as many as ten separate species (Vences et al. 2004; Zimkus 
et al. 2017) with the nominotypical form likely being restricted to Madagascar, the Seychelles, and 
the Mascarenes. While Zimkus et al. (2017) did not include Angolan populations in their molecu¬ 
lar phylogenetic study, it is likely that Angolan populations correspond to both P. nilotica and one 
undescribed species (their OTU 6) that occurs in Gabon and Democratic Republic of Congo. 
Schmidt and Inger (1959) described Ptychadena mascareniensis hylaea from Mount Nimba, 
Liberia and remarked on an individual (CNHM 74214) from “Huila in the highlands of southern 
Angola,” deposited in the Chicago Natural History Museum (now the Field Museum of Natural 
