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PROCEEDINGS OF THE CALIFORNIA ACADEMY OF SCIENCES 
Series 4, Volume 65, Supplement II 
[-17.46777, 23.06667] (Conradie et al. 2016:9-10,18); “Cuando basin (44a)” [-17.57333, 
23.26000] (Conradie et al. 2016:9-10,18). Undetermined Locality: without precise locality 
(Bocage 1870:68; Boulenger 1900:53; Laurent 1954:73); “Rio Quando” (Bocage 1895a: 160; Ruas 
1996:25); “Rio Quanza” (Boulenger 1905:108); “Carangigo” (Boulenger 1882:51); “areas of 
forest and savanna in the north and northeast of Angola” (Cei 1977:16); “plateaus regions” (Cei 
1977:17); “arid territories along the coast” (Cei 1977:17). 
Taxonomic and distributional notes: This species occurs throughout much of sub-Saharan 
Africa in areas of moist savanna, near forest edges, in secondary vegetation with tall herbaceous 
vegetation, and in marshy and agricultura areas (Schmidt and Inger 1959; Ruas 1996; Rodel et al. 
2004; Frost 2016). It appears to be relatively common in Angola, except in the arid and semiarid 
areas in southern regions of the country (Ruas 1996). According to Channing (2001) this species 
often occurs alongside angolensis (Bocage, 1866). 
Ptychadena perpUcata Laurent, 1964 Many-Ridged Grass Frog 
Ptychadena perpUcata Laurent 1964a: 136, fig. 36. Holotype: MD 5513 (collectors A. Barros Machado and 
E. Luna de Carvalho). Type locality: “Alto Chicapa, humidiherhosa des souces du Cuilo” [= Cuilo River] 
Lunda Sul Province, Angola. 
Ptychadena ansorgei: Laurent (1954a:74). 
Ptychadena perpUcata'. Cei (1977:16), Frost (1985:474, 2016), Ruas (1996:27), Largen (2000:177), Channing 
(2001:336). 
Ptychadena {Ptychadena) perpUcata'. Fretey et al. (2011:41). 
Global conservation status (lUCN): Least Concern. 
Global distribution: The species is 
known from Angola and Zambia. 
Ocurrences in Angola (Map 87): Pub¬ 
lished records are limited to the far north of the 
country. Lunda Norte: “Dundo” [-7.36667, 
20.83333] (Laurent 1954a:74,1964a:136; Ruas 
1996:27). Lunda Sul: “Alto Chicapa, humidi- 
herbosa des souces du Cuilo (Rio Cuilo, 
nascente)” [-10.88333, 19.23333] (Laurent 
1964a: 136; Frost 1985:474, 2016; Ruas 
1996:27, Largen 2001:177). 
Taxonomic and distributional notes: 
According to Ruas (1996) and Largen (2000), 
P. perpUcata is only known from forest gal¬ 
leries near the type locality. However, Chan¬ 
ning (2001) provided several records without 
detailed information, presumably based on 
museum specimens, that suggest this species is 
more widespread in the country. 
Ptychadena porosissima (Steindachner, 1867) 
Rana porosissima Steindachner 1867:18, pi. 1, figs. 9-13. Holotype: NHMW 14772 (collector F.A.P. Bayao) 
fide Haupl and Tiedemann (1978:28). Type locality: “Angola”. 
Ptychadena loveridgei Laurent 1954b: 14. Holotype: MRAC 109038. Type locality: “Tare, Busanza, region 
d’Astrida, env. 1. 800 m., Ruanda.” 
i:°E Ifi'E 20°E :4“E 
Map 87. Distribution of Ptychadena perpUcata in 
Angola. 
Striped Grass Frog 
