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Rana porosissima: Bocage (1887a: 191, 1897b:211), Schmidt and Inger (1959:96), Haupl and Tiedemann 
(1978:28), Haupl et al. (1994:32). 
Rana mascareniensis: Bocage (1866a:53), Boulenger (1882:52). 
Rana mascareniensis vw. porosissima'. Bocage (1895a: 160), Ferreira (1897b:240). 
Rana subpunetata: Loveridge (1953a:329). 
Rana (Ptyehadena) maseareniensis: Guibe and Lamotte (1957:978). 
Rana loveridgei: Loveridge (1957:343). 
Ptyehadena porosissima'. Poynton and Broadley (1985b: 149), Frost (1985:474, 2016), Ruas (1996:25), Largen 
(2001:342), Channing (2001:337), Channing et al. (2012:311). 
Ptyehadena {Ptyehadena) porosissima'. Fretey et al. (2011:41). 
Global conservation status (lUCN): Least Concern. 
Global distribution: The species is widely distributed in southern and eastern Africa. 
Ocurrences in Angola (Map 88): The 
species occurs from central-west regions to 
northeastern Angola. Zaire: “S. Salvador do 
Congo” [-6.26667, 14.23333] (Bocage 
1887a:191, Bocage 1895a;160; Ruas 1996:25). 
Kwanza Norte: “Ambaca” [-9.26667, 
15.18333] (Bocage 1895a:160; Ruas 1996:25). 
Malanje: “Duque de Bragan^a” [-9.10000, 
15.95000] (Bocage 1866a:53, 1895a: 160; Bou¬ 
lenger 1882:52; Ruas 1996:25). Lunda Norte: 
“Sombo” [-8.68333, 20.95000] (Laurent 
1954b:14). Benguela: “Quibula” [-12.28333, 
14.68333] (Bocage 1895a:160; Ruas 1996:25); 
“Hanha” [-13.30000, 14.20000] (Bocage 
1897b:211; Ruas 1996:25). Huila: “Rio Cuce 
(Rio Cusse)” [-13.51667, 15.20000] (Ferreira 
1897b:240; Ruas 1996:25); “Caconda” 
[-13.73333, 15.06667] (Bocage 1895a: 160; 
Ferreira 1897b:240; Ruas 1996:25). 
Taxonomic and distributional notes: Ptyehadena porosissima (Steindachner, 1867) was 
considered a variety of Rana mascareniensis (Dumeril and Bibron, 1841) by Bocage (1895a) and 
Ferreira (1897b) and later by Guibe and Lamotte (1957). Ptyehadena loveridgei Laurent, 1954 was 
synonymized with P. porosissima by Schmidt and Inger (1959). Loveridge (1953a) mistakenly 
considered P. porosissima a synonym of P. subpunetata, a position refuted by Schmidt and Inger 
(1959) who considered it a distinct species. Because of the similar general appearance of many 
species of Ptyehadena and the use of characters of limited diagnostic utility in some descriptions, 
many names have been misapplied to more than one species. In this case, some records allocated 
to the P. mascareniensis complex may belong to P. porosissima. 
Map 
Angola. 
Distribution of Ptyehadena porosissima in 
Ptyehadena subpunetata (Bocage, 1866) Spotted Grass Frog 
Rana subpunetata Bocage 1866b:73. Holotype: MBL (collector F.A.P. Bayao) not located by Perret 
(1967a:21), possibly lost or destroyed by fire 18 March 1978. Type locality: “Duque de Braganga” 
[= Calandula] Malanje Province, Angola. 
Rana subpunetata'. Bocage (1866a:54, 1895a: 161, 1897a:203), Inger (1959:541). 
Rana maseareniensis subpunetata'. Mertens (1938a:427). 
Rana subpunetata'. Schmit and Inger (1959:102). 
