140 
PROCEEDINGS OF THE CALIFORNIA ACADEMY OF SCIENCES 
Series 4, Volume 65, Supplement II 
Ptychadena subpunctata: Laurent (1964a: 134), Ferret (1976a:21), Cei (1977:16, 17), Poynton and Broadley 
(19856:143), Frost (1985:476, 2016), Ruas (1996:25, 2002:143), Channing (2001:340), Piekersgill 
(2007a: 136), Conradie et al. (2016:18). 
Ptychadena (Ptychadena) subpunctata'. Fretey et al. (2011:41). 
Global conservation status (lUCN): Least Concern. 
Global distribution: The species is known from Angola, extending east into the southern 
Democratic Republic of Congo and Zambia, and south to northern Botswana and northern 
Namibia. 
Ocurrences in Angola (Map 89): The 
species occurs from central-west to eastern 
areas of Angola. Malanje: “Duque de Bra- 
ganga” [-9.10000, 15.95000] (Bocage 
1866a:54, 1866a:73, 1895a:161, 1897a:203; 
Ferret 1976a;21; Schmidt and Inger 1959:102; 
Poynton and Broadley 1985b: 143; Frost 
1985:476, 2016; Ruas 1996:25). Moxico: 
“Lago Dilolo” [-11.50000, 22.01667] (Ruas 
1996:25, 2002:143); “Lago Cameia” 
[-11.71667, 20.80000] (Ruas 1996:25, 
2002:143); “Rives du lac Calundo” [-11.80000, 
20.86667] (Laurent 1964a: 134; Ruas 1996:25); 
“Chites de la Lusiavo, affl. du Zambeze, 
Calunda” [-11.86667, 23.58333] (Laurent 
1964a: 134; Ruas 1996:25); “Cazombo” 
[-11.88333, 22.91667] (Laurent 1964a:134; 
Ruas 1996:25). Benguela: “Cubal” of Ptychadena subpunctata in 
[-13.03333, 14.25000] (Mertens 1938a:427; 
Ruas 1996:25). Cuando Cubango: “Cubango basin(6b)” [-14.67458, 17.73544] (Conradie et al. 
2016:8-9,18); “Cubango basin (27)” [-15.17127, 19.19433] (Conradie et al. 2016:8-9,18); “Cuito 
basin (30a)” [-17.50875,20.06594] (Conradie et al. 2016:9-10,18); “Cuito basin (30f)” [-17.52638, 
20.05825] (Conradie et al. 2016:9-10,18); “Cuito basin (33a)” [-16.90980, 19.30769] (Conradie et 
al. 2016:9-10,18); “Cuito basin (35)” [-16.62322, 19.05352] (Conradie et al. 2016:9-10,18); 
“Cuando basin (41a)” [-17.46777, 23.06667] (Conradie et al. 2016:9-10,18); “Cuando basin (43)” 
[-17.53500, 23.18916] (Conradie et al. 2016:9-10,18); “Cuando basin (44a)” [-17.57333, 
23.26000] (Conradie et al. 2016:9-10,18); “Cuito basin (56)” [-15.06275, 19.14322] (Conradie et 
al. 2016:9,12,18); “Cuito basin (63)” [-17.93611, 21.10269] (Conradie et al. 2016:9,12,18). Unde¬ 
termined Locality: “areas of forest and savanna in the north and northeast of Angola” (Cei 
1977:16); “plateaus regions” (Cei 1977:16); “Angola” (Poynton and Broadley 1985b: 143); “north 
Angola” (Schmidt and Inger 1959:140). 
Taxonomic and distributional notes: Rana subpunctata was used by Bocage (1866a) as a 
nomen nudum, but in the same issue of the journal, Bocage (1866b) published a valid description 
of the new species. According to Ferret (1976a), the holotype cited by Bocage from “Duque de 
Bragan^a” was not present in the Museu Bocage when he examined that collection. 
