142 
PROCEEDINGS OF THE CALIFORNIA ACADEMY OF SCIENCES 
Series 4, Volume 65, Supplement II 
Ptychadena upembae (Schmidt and Inger, 1959) Upemba Grass Frog 
Rana upembae Schmidt and Inger 1959: 111, fig. 50. Holotype: MRAC 1228 (collector Mission G.F. de Witte), 
formerly Inst. Parcs Natl. Congo Beige fide Lang (1990:13). Type locality: “Kaswabilenga, Parc National 
de I’Upemba, Province Katanga, Belgian Congo” (Schmidt and Inger 1959:111), Democratic Republic of 
Congo. 
Ptyehadena upembae maehadoi Laurent 1964a: 134. Holotype: MD 5364 (unnamed local collector). Type 
locality: “Alto Chicapa, Lunda,’’Angola. 
Ptyehadena upembae maehadoi'. Cei (1977:16). 
Ptyehadena upembae'. Poynton and Broadley (1985b: 150), Frost (1985:477, 2016), Ruas (1996:26, 
2002:144), Channing (2001:342). 
Ptyehadena (Ptyehadena) upembae'. Fretey et al. (2011:42). 
Global conservation status (lUCN): 
Least Concern. 
Global distribution: The species extends 
across south-central African from Angola, the 
southern Democratic Republic of Congo and 
Zambia to Mozambique. 
Ocurrences in Angola (Map 91): The 
species occurs in eastern Angola. Lunda Sul: 
“Alto Chicapa” [-10.88333, 19.23333] (Lau¬ 
rent 1964a: 134; Ruas 1996:25). Moxico: 
“Lago Cameia” [-11.71667, 20.80000] (Ruas 
1996:25, 2002:144). Undetermined Locality: 
“areas of forest and savanna in the north and 
northeast of Angola” (Cei 1977:16). Channing 
(2001) additionnally plotted unspecific locali¬ 
ties in a 2” map. 
Taxonomic and distributional notes: 
Currently P. upembae maehadoi is recognized 
as a synonym of Ptychadena upembae (Channing 
irE I6"E 2(}°E I4'>E 
Map 91. Distribution of Ptychadena upembae in Angola. 
2001; Fretey et al. 2011, Frost 2016). 
Ptychadena uzungwensis (Loveridge, 1932) Udzungwa Grass Frog 
Rana maseareniensis uzungwensis Loveridge 1932a:384. Holotype: MCZ A-16626 (eolleetor A. Loveridge). 
Type loeality: “Dabaga, Uzungwe Mountains, Tanganyika Territory” [Tanzania]. 
Rana maseareniensis subpunetata'. Schmidt (1936:129). 
Ptyehadena uzungwensis'. Laurent (1954b:10, 1964a:139), Cei (1977:16), Poynton and Broadley (1985b:151), 
Poynton and Haacke (1993:14), Frost (1985:477, 2016); Ruas (1996:26, 2002:145), Channing (2001:344^ 
Conradie et al. (2016:18). 
Rana maseareniensis uzungwensis: Loveridge (1953a:372, 1957:342). 
Rana uzungwensis: Schmidt and Inger (1959:117). 
Ptyehadena subpunetata: Ruas (1996:25). 
Ptyehadena (Ptyehadena) uzungwensis: Fretey et al. (2011:42). 
Global conservation status (lUCN): Least Concern. 
Global distribution: The species is known from central eastern Angola and the southeastern 
Democratic Republic of Congo, Zambia and Tanzania south to northern Zimbabwe. 
Ocurrences in Angola (Map 92): The species occurs chiefly in eastern Angola. Lunda 
Norte: “Dundo” [-7.36667, 20.83333] (Laurent 1954b: 10). Lunda Sul: “Alto Chicapa, hemidi- 
