MARQUES ET AL.; AMPHIBIANS AND TERRESTRIAL REPTILES OF ANGOLA 
143 
herbosa pres de la cascade du Cuango-Muque” 
[-10.76667, 19.20000] (Laurent 1964a:139); 
“Alto Chicapa, Lunda hemidiherbosa des sour¬ 
ces du Cullo” [-10.86667, 19.40000] (Laurent 
1964a: 139); “Alto Chicapa” [-10.88333, 
19.23333] (Laurent 1964a:139); “Dala” 
[-11.03333, 20.20000] (Laurent 1964a:139). 
Malanje: “20 km NW of Quimbango” 
[-10.76667, 17.43333] (Poynton and Haacke 
1993:14). Mexico: “Calombe” [-11.83333, 
19.93333] (Ruas 2002:145); “Reserve de chas- 
se de Cameia dans la steppe de Narikumbi” 
[-11.83333, 21.00000] (Laurent 1964a:139). 
Bie: “Gauca” [-11.18333, 17.45000] (Schmidt 
1936:129; Laurent 1954b:9; Schmidt and Inger 
1959:118); “Chitau” [-11.43333, 17.15000] 
(Schmidt 1936:129; Laurent 1954b:9; Schmidt 
and Inger 1959:118). Cuando Cubango: 
“Cuito basin (27)” [-15.17127, 19.19433] 
(Conradie et al. 2016:8-9,18). Undetermined Locality: “areas of forest and savanna in the north 
and northeast of Angola”(Cei 1977:16). 
Taxonomic and distributional notes: Loveridge (1953a) believed thaX Rana mascareniensis 
uzungwensis Loveridge, 1932 was erroneously synonymized with Rana subpunctata (Bocage, 
1866) by Schmidt (1936) and cited specimens from “Gauca” and “Chitau” in Angola as belonging 
to P. uzungwensis. This species is considered valid by the majority of modem authors (e.g., Poyn¬ 
ton and Broadley 1985b; Channing 2001; Frost 2016) and according to Poynton and Broadley 
(1985b) it inhabits grasslands and dambos. Poynton and Haacke (1993) reported one individual 
from “20 km NW of Quimbango” collected by G. G. A. Voigt in a water-logged grassland. Chan¬ 
ning (2001) provided several records in the western regions of the country based on previously 
unpublished museum records. 
Family Phrynobatrachidae Laurent, 1941 
Genus Phrynobatrachus Gunther, 1862 
Phrynobatrachus brevipalmatus (Ahl, 1923) Ahl’s Screeching Frog (Endemic) 
Hylarthroleptis brevipalmatus Ahl “1923” 1925:102. Holotype: ZMB 26689 (collector K. May). Type local¬ 
ity: “Loanda” [= Luanda] Luanda Province, Angola. 
Phrynobatrachus brevipalmatus'. Fretey et al. (2011:38), Frost (2016). 
Global conservation status (lUCN): Data Deficient. 
Global distribution: The species is only known from Angola. 
Ocurrences in Angola (Map 93): The species is known from Luanda. Luanda: “Loanda” 
[-8.83333, 13.26667] (Ahl “1923” 1925:102; Frost 2016). 
Taxonomic and distributional notes: According to the original description, this species 
is closely related to Hylarthroleptis graueri (Nieden, 1911) from eastern Africa. This species was 
not discussed by Zimkus et al. (2010) and no further details are available for its biology and 
systematics. 
1I°E I6'E irE 24°E 
Map 92. Distribution of Ptychadena uzungwensis in 
Angola. 
