MARQUES ET AL.; AMPHIBIANS AND TERRESTRIAL REPTILES OF ANGOLA 
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17.73544] (Conradie et al. 2016:8-9,16); “Cuando basin (19)” [-14.70213, 17.37772] (Conradie et 
al. 2016:8-9,16); “Cuando basin (21)” [-14.66586,17.07661] (Conradie et al. 2016:8-9,16); “Cuan¬ 
do basin (22e)” [-14.65386, 16.93547] (Conradie et al. 2016:8-9,16); “Cuando basin (22d)” 
[-14.64991, 16.90739] (Conradie et al. 2016:8-9,16); “Cuando basin (29)” [-17.87291, 19.833333] 
(Conradie et al. 2016:9-10,16); “Cuito basin (35)” [-16.62322, 19.05352] (Conradie et al. 2016: 
9-10,16); “Cuando basin (38)” [-17.58830, 22.65694] (Conradie et al. 2016:9-10,16); “Cuando 
basin (39)” [-17.46333, 22.86638] (Conradie et al. 2016:9-10,16); “Cuando basin (40)” 
[-17.45786, 22.91191] (Conradie et al. 2016:16); “Cuando basin (43)” [-17.53500, 23.18916] 
(Conradie et al. 2016:9-10,16); “Cuando basin (44a)” [-17.57333, 23.26000] (Conradie et al. 
2016:9-10,16). Phrynobatrachus minutus (plotted as pentagons). Lunda Norte: “Dundo” 
[-7.36667, 20.83333] (Laurent 1954a:74; Ruas 1996:27); “galerie forestiere de la Luaehimo (Lua- 
chimo)” [-7.38333, 20.85000] (Laurent 1950a: 15; Ruas 1996:27); “Marrura rive droite de la Tshi- 
kapa, 50 km sud-oest de Dundo (Marrora)” [- 7.60000, 20.51667] (Laurent 1954a:74; Ruas 
1996:27). Cunene: “Donguena (Cunene)” [-17.01667, 14.71667] (Laurent 1954a:74; Ruas 
1996:27). Undetermined Locality: “Station Quanza (Esta^ao do Quanza)” (Laurent 1954a:74; 
Ruas 1996:27); with no preeise loeality (Laurent 1954a:74); “areas of forest and savanna in the 
north and northeast of Angola” (Cei 1977:16); “plateaus regions” (Cei 1977:17). Phrynobatrachus 
parvulus (plotted as diamonds). Lunda Sul: “Alto Cuilo, Poste de Caeolo, ruisseau cascade Na- 
Ipanha” [-10.00000, 19.58333] (Laurent 1964a:144); “Alto Cuilo, rives du Cuilo (Cuilo)” 
[-10.01667, 19.55000] (Laurent 1964a: 144); “Alto Chicapa, humidiherbosa des sources de la 
Kamutongola” [-10.88333, 19.25000] (Laurent 1964a: 144). Moxico: “Teixeira de Sousa” 
[-10.70000, 22.23333] Mertens (1937:20). Malanje: “Range N’golla (Dange)” [-8.43333, 
16.56667] (Boulenger 1905:109; Loveridge 1933:386; Marx 1958:425; Schmidt and Inger 
1959:160; Poynton and Broadley 1985b:169; Frost 1985:449, 2016; Ruas 1996:27). Kwanza Sul: 
“Calaongo below Congulu (Carlaongo)” [-10.78333, 14.26667] (Parker 1936:142; Ruas 1996:27). 
Bie: “Gauca” [-11.18333, 17.45000] (Schmidt 1936:131; Ruas 1996:27); “Chitau” [-11.43333, 
17.15000] (Schmidt 1936:131, Ruas 1996:27); “General Machado” [-12.03333,17.46667] Mertens 
(1937:20). Huambo: “Bimbi” [-11.81667, 15.83333] (Monard 1937a:58, 1938:118, Ruas 
1996:27). Benguela: “Cubal” [-13.03333, 14.25000] Mertens (1937:20). Cuando Cubango: 
“Cuando basin (43)” [-17.53500, 23.189916] (Conradie et al. 2016:9-10,17). Undetermined 
Locality: “without precise locality” (Schmidt 1936:131). 
Taxonomic and distributional notes: Opinions in the literature vary as to whether three 
commonly reported species from Angola are conspecific: P. mababiensis FitzSimons, 1932, 
P. parvulus (Boulenger, 1905) and P. minutus (Boulenger, 1895). Loveridge (1953) assigned a 
specimen previously identified as P. parvulus to P. mababiensis, suggesting that P. parvulus was a 
western relative of P. mababiensis or perhaps a synonym of Phrynobatrachus minutus. Poynton 
and Broadley (1985b:165, 170) discussed difficulties in separating P. parvulus and P. mababiensis 
based on external morphology. Zimkus et al. (2010) recently used molecular phylogenetic analy¬ 
ses to reveal that there are probably three different species currently recognized as P. mababiensis. 
If P. minutus is known only from Ethiopia, it is highly likely that the Angolan records assigned to 
this species are misidentified and instead belong to one of the other three species. With some reser¬ 
vations, Laurent (1964a) cited P. cryptotis for Angola, though he recognized that these might 
represent P. parvulus instead, which is a similar species (Drewes and Vindum 1994). It is likely that 
the Angolan specimens mapped here represent a combination of P. mababiensis and 
P. parvulus. 
