MARQUES ET AL.; AMPHIBIANS AND TERRESTRIAL REPTILES OF ANGOLA 
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Phrynobatrachus natalensis (Smith, 1849) Natal Dwarf Puddle Frog 
Stenhorhynchus natalensis Smith 1849b:24. Holotype: BMNH 1947.2.5.13 (formerly BMNH 62.3.14.20) 
(collector A. Smith) fide Grandison in Frost (1985:448). Type locality: “the country around Port Natal” 
[= Durban, KwaZulu-Natal], South Africa. 
Phrynobatraehus natalensis: Gunther (1865a:481), Bocage (1866a:54, 1870:68, 1879c:89, 1895a:162, 
1897b:211), Boulenger (1882:112, 1905:108), Ferreira (1904:113, 1906:166), de Witte (1919:223), 
Sc hmi dt (1936:130), Monard (1937a:57, 1938:117), Mertens (1937a:20), Parker (1939:142), Laurent 
(1950a: 15, 1954a:74, 1964a: 143), Loveridge (1957:348), Hellmich (1957a:24), Schmidt and Inger 
(1959:155), Inger (1959:540), Cei (1977:16, 17, 18), Poynton and Haacke (1993:14), Ruas (1996:27, 
2002:145), Largen (2001:348), Channing (2001:307), Zimkus et al. (2010:886), Fretey et al. (2011:39), 
Frost (2016), Ceriaco et al. (2016b:39), Conradie et al. (2016:17). 
Phrynobatraehus cf natalensis: Ceriaco et al. (2014b:669). 
Global conservation status (lUCN): Least Concern. 
Global distribution: Populations referred to P. natalensis occur across sub-Saharan Africa, 
including in Angola, though this taxon likely contains multiple undescribed species. 
Ocurrences in Angola (Map 95): The 
species is widespread across nearly all of 
Angola. Lunda Norte: “Dundo” [-7.36667, 
20.83333] (Laurent 1950a; 15, 1954a:74; Ruas 
1996:27); “Andrada (Luembe O)” [-7.70000, 
21.38333] (Laurent 1954a;74; Ruas 1996:27); 
“Muita (Luembe E)” [-7.80000, 21.45000] 
(Laurent 1950a: 15, 1954a:74; Ruas 1996:27). 
Lunda Sul: “Dala” [-11.03333, 20.20000] 
(Monard 1937a:57, 1938:117; Ruas 1996:27); 
“Alto Cuilo, rives du Cuilo (rio Cuilo)” 
[-10.01667, 19.55000] (Laurent 1964a:143; 
Ruas 1996:27). Malanje: “Bange N’golla 
(Dange)” [-8.43333, 16.56667] (Boulenger 
1905:108; Ruas 1996:27); “Duque de Bragan- 
ga” [-9.10000, 15.95000] (Bocage 1866a:54, 
1895a: 162; Gunther 1865a:481; Boulenger 
1882:112; de Witte 1919:223; Poynton and 
Haacke 1993:14; Ruas 1996:27); “16 km SE of 
Quissol - Garibo” [-9.63333, 16.41667] (Poynton and Haacke 1993:14; Ruas 1996:27); “Cangan- 
dala National Park” [-9.84606, 16.72233] (Ceriaco et al. 2016b:39); “Capanda” [-11.58333, 
17.61667] (Ceriaco et al. 2014b:669); “ Reserva da Palanca Preta (nascentes do rio Cuanza)” 
[-11.11667, 17.46667] (Ruas 2002:145); “Rio Caluando (nascente)” [-11.11667, 17.46667] (Ruas 
1996:27, 2002:145). Mexico: “Teixeira de Sousa” [-10.70000, 22.23333] (Mertens 1937:20); 
“Posto Bussaco” [-11.43333, 20.16667] (Ruas 2002:145; Ruas 1996:27); “Rives du lac Calundo 
(Lago Calundo)” [-11.80000, 20.86667] (Laurent 1964a: 143; Ruas 1996:27); “Rio Calombe 
(Reserva da Palanca Preta)” [-11.83333, 19.93333] (Ruas 1996:27, 2002:145); “Cazombo” 
[-11.88333, 22.91667] (Laurent 1964a:143; Ruas 1996:27). Kwanza Norte: “Mucoso” 
[-9.53333, 14.65000] (Hellmich 1957a:24). Bengo: “Catete” [-9.11667, 13.70000] (Ferreira 
1904:113; Ruas 1996:27). Kwanza Sul: “Congulu (Congulo)” [-10.86667, 14.28333] (Parker 
1936:142; Ruas 1996:27). Bie: “Gauca” [-11.18333, 17.45000] (Schmidt 1936:130; 
Ruas 1996:27); “Chitau” [-11.43333, 17.15000] (Schmidt 1936:130; Ruas 1996:27); “General 
Map 95. Distribution of Phrynobatrachus natalensis in 
Angola. 
