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PROCEEDINGS OF THE CALIFORNIA ACADEMY OF SCIENCES 
Series 4, Volume 65, Supplement II 
Maehado” [-12.03333, 17.46667] (Mertens 1937a:20); “Bihe” [-12.38333, 16.95000] (Bocage 
1879a:89, 1895a: 162; de Witte 1919:223; Ruas 1996:27); “Cubango basin (10)” [-13.71616, 
17.09661] (Conradie et al. 2016:8-9,17); “Cubango basin (12a)” [-13.59333, 16.87986] (Conradie 
et al. 2016:17); “Cubango basin (17)” [-12.57008, 16.49111] (Conradie et al. 2016:8-9,17). Huam- 
bo: “Bimbi” [-11.81667, 15.83333] (Monard 1937a:57, 1938:117; Ruas 1996:27). Benguela: 
“Quissange” [-12.43333, 14.05000] (Boeage 1895a:162; de Witte 1919:223; Ruas 1996:27); 
“Quindumbo” [-12.46667, 14.93333] (Boeage 1895a:162; de Witte 1919:223; Ruas 1996:27); 
“Ebanga” [-12.73333, 14.73333] (Monard 1937a:57, 1938:117; Ruas 1996:27); “Entre Rios” 
[-13.01667, 14.63333] (Hellmich 1957a:24); “Hanha” [-13.30000, 14.20000] (Bocage 1897b:211; 
Ruas 1996:27). Hufla: “Caconda” [-13.73333, 15.06667] (Bocage 1895a:162; de Witte 1919:223; 
Ruas 1996:27); “Kalukembe” [-13.78333, 14.68333] (Monard 1937a:57, 1938:117; Ruas 
1996:27); “Sangeve” [-13.88333,15.83333] (Monard 1937a:57, 1938:117; Ruas 1996:27); “Muko- 
ti” [-14.20000, 15.80000] (Monard 1937a:57, 1938:117; Ruas 1996:27); “Kuvangu” 
[-14.46667, 16.30000] (Monard 1937a:57, 1938:117; Ruas 1996:27); “Boca de Humpata” 
[-14.93333, 13.51667] (Laurent 1964a:143; Ruas 1996:27); “Kangela (Kului)” [-15.41667, 
15.73333] (Monard 1937a:57; 1938:117; Ruas 1996:27). Namibe: “Cunene mouth (Foz do Cune- 
ne)” [-17.28333, 11.80000] (Poynton and Haacke 1993:14; Ruas 1996:27). Cunene: “ruisseau 
Mbale (Bale)” [-15.16667, 16.75000] (Monard 1937a:57, 1938:117; Ruas 1996:27); “Kuvelai” 
[-15.65000, 15.80000] (Monard 1937a:57, 1938:117; Ruas 1996:27); “Donguena” 
[-17.01667, 14.71667] (Laurent 1954a:74; Ruas 1996:27). Cuando Cubango: “Kakindo” 
[-15.45000, 17.05000] (Monard 1938:117; Ruas 1996:27); “Cubango basin (3)” [-14.94277, 
17.71863] (Conradie et al. 2016:8-9,17); “Cubango basin (6b)” [-14.67458, 17.73544] (Conradie 
et al. 2016:8-9,17); “Cubango basin (19)” [-14.70213, 17.37772] (Conradie et al. 2016:8-9,17); 
“Cubango basin (21)” [-14.66586, 17.07661] (Conradie et al. 2016:17); “Cubango basin (22d)” 
[-14.66278, 16.96081] (Conradie et al. 2016:17). Undetermined Locality: without precise locali¬ 
ty (Bocage 1870:68; Ferreira 1906:166; Schmidt 1936:130); “areas of forest and savanna in the 
north and northeast of Angola” (Cei 1977:16); “plateaus regions” (Cei 1977:17); “arid territories 
along the coast” (Cei 1977:18). 
Taxonomic and distributional notes: Schmidt and Inger (1959) provided a distribution map 
for Phrynobatrachus natalensis (Smith, 1849) with some records in Angola, though without addi¬ 
tional information. The molecular phylogenetic analysis by Zimkus et al. (2010) VQYQdlQd Phryno¬ 
batrachus natalensis to be a species complex, with Angolan records likely corresponding to their 
lineage ''natalensis A.” 
Phrynobatrachus plicatus (Gunther, 1858) Coast River Frog 
Hyperolius plicatus Gunther 1858a:326. Holotype: BMNH 1947.2.29.97 [wrongly given as 1947.2.29.47 by 
Grandison in Frost (1985)], formerly BMNH 49.10.9.1 fide Grandison in Frost (1985:449) (eolleetor 
unknown). Holotype is depleted in Gunther ‘T858b” 1859:8, pi. 7, fig. C. Type loeality: “Guinea” (Giin- 
ther 1858a:326), later modified to “Coast of Guinea” (Gunther ‘T858b” 1859:88). 
Arthroleptis plicatus: Peters (1877a:618). 
Phrynobatrachus plicatus: Zimkus et al. (2010:896), lUCN SSC Amphibian Speeialist Group (2013e), Frost 
(2016). 
Global conservation status (lUCN): Least Concern. 
Global distribution: This is a widespread species from western Africa. 
Ocurrences in Angola (Map 96): The only record of this species in Angola is from the Cabin¬ 
da enclave. Cabinda: “Chinchoxo” [-5.10000, 12.10000] (Peters 1877a:618). 
Taxonomic and distributional notes: The record of this species in Angola is almost certain- 
