MARQUES ET AL.: AMPHIBIANS AND TERRESTRIAL REPTILES OF ANGOLA 
I5I 
[-13.45000,14.61667] (Poynton and Haacke 1993:14; Ruas 1996:23); “Tongrube am Jamba-FluP” 
[-13.60000, 16.60000] (Hellmich I957a:25); “Caconda” [-13.73333, 15.06667] (Bocage 
I895a:I58; Perret 1976:18; Gavetti and Andreone 1993:91; Ruas 1996:23); “Kalukembe” 
[-13.78333, 14.68333] (Monard I937a:43, 1938:99; Ruas 1996:23); “Sangeve” [-13.88333, 
15.83333] (Monard I937a:43, 1938:99; Ruas 1996:23); “Kuvangu” [-14.46667, 16.30000] 
(Monard I937a:43, 1938:99; Ruas 1996:23); “Boca de Humpata” [-15.01667, 13.38333] (Laurent 
1964a: 132; Ruas 1996:23; Channing et al. 2012:318; Channing and Baptista 2013:508); “Zootec- 
nica Station, Humapata” [-I4.9I4I7, 13.31653] (Channing et al. 2016:16, 66); “Humapata eamp” 
[-14.95000, 13.26667] (Channing et al. 2016:16, 66); “Huila” [-15.05000, 13.55000] (Boeage 
1895a:158; Themido 1941:2; Perret 1976a:18; Ruas 1996:23). Namibe: “Lebapass, between river 
and highway” [-15.07006, 13.24414] (Ceriaeo et al. 2016a:20). Cuando Cubango: “Cubango 
basin (4)” [-14.81913, 17.67450] (Conradie et al. 2016:18); “Cubango basin (22d)” [-14.64991, 
16.90739] (Conradie et al. 2016:18); “Cuando Cubango Provinee” [-14.64991, 18.4672] (Chan¬ 
ning et al. 2016:16). Undetermined Locality: “without precise locality” (Bocage 1870:78); “Rio 
Quando” (Bocage 1895a: 158; Ruas 1996:23); “areas of forest and savanna in the north and north¬ 
east of Angola” (Cei 1977:16); “plateaus regions” (Cei 1977:16); “arid territories along the coast” 
(Cei 1977:16); “Mombola” (Channing et al. 2016:16). 
Taxonomic and distributional notes: Many authors, including Poynton (1964), Drewes and 
Vindum (1994), and Channing and Howell (2006) recognized that there are probably several unde¬ 
scribed cryptic species within the enormous range of this taxon. Pickersgill (2007a) named three 
new species from Eastern Africa populations that were previously included within Amietia 
angolensis. Channing and Baptista (2013) and Channing et al. (2016) restricted nominal 
A. angolensis to Angolan populations, though recognized that it may occur more broadly, espe¬ 
cially in neighboring countries for which they lacked specimens with genetic data. 
Genus Boulenger, 1917 
Aubria sp. 
Rana subsigillata Dumeril 1856:560. Holotype: MNHN 1566 (collector J. Aubry-Lecomte). Type locality: 
“Gabon” (Dumeril 1856:560). 
Aubria masako Ohler and Kazadi 1990:29, figs. 4, 8, 10, 12, 24. Holotype: MNHN 1989.2775 (collector 
M. Kazadi). Type locality: “Foret de Masako pres du village Batiabongena a 15 km du centre-ville de 
Kisangani sur I’ancienne route Buta (Zaire)” (Ohler and Kazadi 1990:29), [= Masako forest, near 
Batiabongena village, 15 km from Kisangani], Democratic Republic of Congo. 
Rana {Aubria) subsigillata: Monard (1937a:47, 1938:104). 
Aubria subsigillata: Cei (1977:17), Perret (1996:96). 
Aubria masako: Channing (2011:284), Fretey et al. (2011:42), Channing et al. (2012:329), Frost (2016). 
Global distribution: Aubria masako and A. subsigillata are known from the tropical forests 
of central Africa, with ranges that together extend from the Atlantic coast of Cameroon and Gabon 
and into the Congo Basin of Democratic Republic of Congo. 
Ocurrences in Angola (Map 98): Both Aubria subsigillata and A. masako have only been 
reported from “Kakindo (Kuvangu)” in southern Angola. Cuando Cubango: “Kakindo (Kuvan¬ 
gu)” [-14.46667, 16.30000] (Monard 1937a:47, 1938:104; Channing 2001:285). 
Taxonomy and natural history notes: Monard (1937a, 1938) cited one specimen of Aubria 
subsigillata (Dumeril, 1856) from “Kakindo (Kuvangu),” Angola that was identified by Gaston de 
Witte. This first and only record for the genus in Angola is unlikely due to the habitat, a dry savan¬ 
na, and its long distance from congeneric populations in central or western Africa. Perret (1996) 
revisited this specimen and cited morphological differences from A. subsigillata in Cameroon to 
