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PROCEEDINGS OF THE CALIFORNIA ACADEMY OF SCIENCES 
Series 4, Volume 65, Supplement II 
suggest that it might represent a new species or 
even a new genus. It remains unclear whether 
this record indeed corresponds to Aubria and, if 
so, to which species it should be referred. The 
specimen cited by Monard (1937a, 1938) was 
recently located in the collections of the Musee 
d’Histoire Naturelle, La-Chaux-de-Fond, 
Switzerland and is presently being studied 
(Ceriaco et al. in prep.). 
12°E l(j®E 2()°E 24'*E 
Map 98. Distribution oi Aubria sp. in Angola. 
Genus Pyxicephalus Tschudi, 1838 
Pyxicephalus edulis Peters, 1854 Edible Bullfrog 
Pyxicephalus edulis Peters 1854:626. Leetotype: ZMB 50301 (formerly part of ZMB 10056) (eolleetor: 
W.C.H. Peters) designated by Seott et al. (2013:201). Type loeality: restrieted to “Tete,” Mozambique by 
Loveridge (1953:375). The original syntype series ineluded ZMB 3349, 3350 [4 speeimens from Mozam¬ 
bique], 10056 [2 speeimens (Tete)], and 10057 [2 speeimens (Sena)]. Bauer et al. (1995:49) mentioned 
ZMB 10058 (not loeated), 50260, 50290, and 50301-02, formerly ineluded under the earlier numbers, one 
of these may be the BMNH speeimen from “Mossambique” mentioned by Boulenger (1882:34) as “typi- 
eal of P. edulis"' and reeeived from W. Peters. Original type locality: “Mosambique, Boror, Tette” (Peters 
1854:626), [= Mozambique Island and adjacent mainland, Boror, Tete] 
Rana adspersa: Bocage (1895a: 157), Inger (1959:541). 
Rana (Pyxieephalus) adspersus: Monard (1937a:46, 1938:103), Frade (1963:254). 
Pyxieephalus adspersus: Cei (1977:17), Frost (1985:477), Channing (2001:346), du Preez and Camthers 
(2009:414), Channing et al. (2012:123). 
Pyxieephalus adspersus edulis: Loveridge (1950:255, 1953a:375), Parry (1982:286), Poynton and Broadley 
(1895b: 123), Poynton and Haacke (1993:13), Ruas (1996:23). 
Pyxieephalus edulis: Channing et al. (1994:154), Fretey et al. (2011:42), Scott et al. (2013:201), Frost (2016). 
Global conservation status (lUCN): Least Concern. 
Global distribution: The species is broadly distributed in sub-Saharan Africa, extending from 
Senegal to Kenya, and south through Mozambique, Malawi, Angola, into easternmost Namibia and 
northeastern South Africa. 
Ocurrences in Angola (Map 99): The species is known from southern Angola. Cunene: 
“Humbe” [-16.68333, 14.90000] (Bocage 1895a:157; Ruas 1996:23); “23 km NW of Pereira de 
Eqa (Roqadas)” [-16.95000, 15.56667] (Parry 1982:286; Poynton and Haacke 1993:13; Ruas 
1996:23); “Pereira de Eqa” [-17.06667, 15.73333] (Poynton and Haacke 1993:13; Ruas 1996:23); 
“Mupanda” [-17.13333, 15.76667] (Monard 1937a:46, 1938:103; Ruas 1996:23). 
Taxonomic and distributional notes: There has been on-going confusion about the taxono¬ 
my of Pyxicephalus edulis Peters 1854 andR adspersus Tschudi 1838. The former was recognized 
as a subspecies of the latter by Boulenger (1882) and many subsequent authors (Loveridge 1950, 
