MARQUES ET AL.; AMPHIBIANS AND TERRESTRIAL REPTILES OF ANGOLA 
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1953a; Parry 1982; Poynton and Broadley 
1895b; Poynton and Haacke 1993; Ruas 1996), 
though Channing et al. (1994) elevated 
P. edulis based on differenees in the ealls and 
breeding biology, as well as some morphologi- 
eal differenees (Seott et al. 2013). Channing et 
al. (1994) also suggested that much of the liter¬ 
ature regarding Pyxicephalus adspersus should 
instead apply to P. edulis. Several authors 
including Channing (2001), du Preez and 
Caruthers (2009), and Channing et al. (2012) 
considered that P. edulis does not occur in 
Angola, whereas Fretey et al. (2011) consid¬ 
ered it present there. Ceriaco et al. (in prep.) 
recently examined Monard’s (1937a, 1938) 
specimens from “Mupanda” identified as Rana 
{Pyxicephalus) adspersus in the Musee d’his- 
toire naturelle de La Chaux-de-Fonds, Switzer- Distribution of Pyxicephalus edulis in Angola. 
land and considered that these should be referred to P. edulis based on distinctive morphological 
characters. 
i:°E lO'E 2'0°E 24*E 
Genus Tomopterna Dumeril and Bibron, 1841 
Tomopterna cryptotis (Boulenger, 1907) Tremelo Sand Frog 
Rana cryptotis'QouXengex 1907b:109. Syntypes: BMNH 1947.2.1.73-78, 1947.2.28.48-5, and MCZ A-19268, 
exchanged from BMNH fide Barbour and Loveridge (1946:182) (collector Dr. W.J. Ansorge), formerly 
BMNH 1907.6.29.82-96. Type locality: “Catequero,” “Ponang Kuma (Dongwenna)” and “Kafitu 
swamps” (Boulenger 1907b: 109) Namibe Province, Angola. 
Rana cryptotis'. Barbour and Loveridge (1946:182). 
Tomopterna cryptotis: Poynton and Broadley (1985b: 125), Poynton and Haacke (1993:13), Frost (1985:528, 
2016), Ruas (1996:23), Channing (2001:365), Largen (2001:330), Channing and Howell (2006:329), Pick- 
ersgill (2007a: 118). 
Tomopterna {Tomopterna) cryptotis: Fretey et al. (2011:43). 
Tomopterna cryptotis: Conradie et al. (2016:19). 
Global conservation status (lUCN): Least Concern. 
Global distribution: This widespread species is found in xeric regions across much of sub- 
Saharan Africa. 
Ocurrences in Angola (Map 100): The species occurs in southwestern Angola. Namibe: “25 
km W of Virei” [-15.66667, 12.76667] (Poynton and Haacke 1993:13; Ruas 1996:23); “Miranda” 
(Boulenger 1907b: 109; Poynton and Haacke 1993:13; Ruas 1996:23). Cunene: “2 km NW of 
Calequero (Cahama)” [-16.28333, 14.30000] (Poynton and Haacke 1993:13); “Catequero” 
[-16.56667, 14.90000] (Boulenger 1907b:109; Poynton and Broadley 1985b:125; Frost 2016, 
1985:528; Ruas 1996:23; Largen 2001:330); “Ponang Kuma (Dongwenna)” [-17.05000, 14.65000] 
(Boulenger 1907b: 109; Barbour and Loveridge 1946:182; Poynton and Broadley 1985b: 125; Frost 
2016, 1985:528; Ruas 1996:23); “Kafitu swamps” (Boulenger 1907b: 109; Poynton and Broadley 
1985b: 125; Frost 2016, 1985:528; Largen 2001:330). Cuando Cubango: “Cubango basin (22c)” 
[-14.65386, 16.93547] (Conradie et al. 2016:8-9,19); “Cubango basin (29)” [-17.87291, 19.83333] 
(Conradie et al. 2016:9-10,19). 
