154 
PROCEEDINGS OF THE CALIFORNIA ACADEMY OF SCIENCES 
Series 4, Volume 65, Supplement II 
Taxonomic and distributional notes: 
Originally described based on specimens from 
Angola, many subsequent authors have identi¬ 
fied populations across much of Africa as 
T. cryptotis. Recent molecular genetic analyses 
suggest at least some populations found far 
from Angola, including Somalia and Kenya, 
represent other distinct species (Zimkus and 
Larson 2011; Wasonga and Channing 2013). 
Because of the difficulty in identifying some 
species of Tomopterna based on morphology 
alone, the extent of the distribution of this 
species remains uncertain (Channing 2001; 
Channing and Howell 2006). Some records of 
T. cryptotis might refer instead to T. tandyi. 
Channing (2001) provided a map for T. tandyi 
with records from southern Angola (Pickersgill 
2007a; Channing et al. 2012), between Namibe 
and Cunene provinces. 
12°E l(j®E 2i>°E 24'*E 
Map too. Distribution of Tomopterna cryptotis in 
Angola. 
Tomopterna damarensis Dawood and Channing, 2002 Damaraland Sand Frog 
Tomopterna damarensis Dawood and Channing 2002:130, figs 1, 2, 3. Holotype: TM 83913 (collector 
A. Channing). Type locality: “Khorixas” (Dawood and Channing 2002:133), Kunene Region, Namibia. 
Tomopterna damarensis: Frost (2016), Ceriaco et al. (2016a:20). 
Global conservation status (lUCN): 
Data Deficient. 
Global distribution: The species is 
known from southwestern Angola and northern 
Namibia. 
Ocurrences in Angola (Map 101): The 
species occurs in southwestern Angola. 
Namibe: “Pediva Hot springs” [-16.28462, 
12.56329] (Ceriaco et al. 2016a;20). 
Taxonomic and distributional notes: 
Dawood and Channing (2002) suggested that 
this species might occur more broadly. Recent¬ 
ly Ceriaco et al. (2016a) reported the first 
record for Angola from Iona Natinoal Park. 
Previous records of some other species of 
Tomopterna, including T. tandyi, might refer to 
this species (Heinicke et al. 2017). 
1I°E It'E 20°E 24”E 
Map 101. Distribution of Tomopterna damarensis in 
Angola. 
Tomopterna krugerensis Passmore and Carruthers, 1975 Knocking Sand Frog 
Tomopterna krugerensis Passmore and Carmthers 1975:32, figs. 4,5,6,7, 8. Holotype: TM 44670 (collectors 
N.I. Passmore and V.C. Carmthers). Type locality: “Machayipan, Kmger National Park,” South Africa. 
