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Tomopterna krugerensis: Poynton and Broadley (1985b: 127), Channing (2001:368). 
Tomopterna {Tomopterna) krugerensis: Fretey et al. (2011:43). 
Global conservation status (lUCN): Least Concern. 
Global distribution: The species is occurs in savannas extending from Angola and Namibia, 
east through Botswana into northern South African and southern Mozambique. 
Ocurrences in Angola: The species occurs in southwestern Angola. Undetermined Locali¬ 
ty: without precise locality (Channing 2001:369). 
Taxonomic and distributional notes: This species is similar morphologically to T. cryptotis 
but has a distinct call (Poynton and Broadley 1985b) and is phylogenetically distinct (Zimkus and 
Larson 2011). Because of the morphological similarity, some previous reports from Angola for 
T. cryptotis might be referable to this species or T. tandyi. Channing (2001) presented the only 
records in Angola for T. krugerensis with mapped points in Namibe and Cunene provinces, though 
without specific locality details. 
Tomopterna tandyi Channing and Bogart, 1996 Tandy’s Sand Frog 
Tomopterna tandyi Channing and Bogart 1996:80, figs. 2, 3. Holotype: PEM A-2283 (collector M. Snyman). 
Type locality: “Bedford in the Eastern Cape” (Channing and Bogart 1996:80), South Africa. 
Tomopterna tandyi: Channing (2001:372), Pickersgill (2007a: 117), Fretey et al. (2011:43), Channing et al. 
(2012:360), Frost (2016). 
Global conservation status (lUCN): Least Concern. 
Global distribution: The species occurs in South Africa, Namibia, and southwestern Angola. 
Ocurrences in Angola: The species occurs in southwestern Angola. Undetermined Locali¬ 
ty: “without precise locality” (Channing 2001:369). 
Taxonomic and distributional notes: A tetraploid species, likely of hybrid origin between 
Tomopterna cryptotis (Boulenger, 1907) and Tomopterna delalandii (Tschudi, 1838) (Channing 
and Bogart 1996, Frost 2016). This species is morphologically indistinguishable from T. crypto¬ 
tis and T. delalandii and many records of Tomopterna cryptotis, including in Angola, may refer to 
T. tandyi. Channing (2001) provided a map for T. tandyi with records from southwestern Angola 
between Namibe and Cunene provinces (followed by Pickersgill 2007a, Channing et al. 2012, and 
Frost 2016), though without specific information. 
Tomopterna tuberculosa (Boulenger, 1882) Rough Sand Frog 
Pyxieephalus rugosus Gunther 1865a:479, pi. 33, fig. 1. Syntypes: BMNH 1947.2.29.10-11 (collector F.M.J. 
Welwitsch), formerly BMNH 64.7.13.5-6. Type locality: “Pungo Andongo” and “W. Africa” (Gunther 
1865a:479), Angola. Preoccupied by Rana rugosa Temminck and Schlegel, 1838. 
Rana tuberculosa Boulenger 1882:30. Replacement name for Pyxieephalus rugosus Gunther, 1865. 
Rana {Tomopterna) signata A\\\ “1923” 1925:43. Syntypes: ZMB 6458, 7765 [2 specimens] (donor J.V.B. 
Bocage). Type locality: “Huilla” [= Huila], Malanje Province, Angola. Synonymy by Laurent (1954a:72). 
Rana {Tomopterna) eaeondana Ahl “1923” 1925:43. Holotype: ZMB 15477 (purchased Linnea). Type local¬ 
ity: “Caconda,” Huila Province, Angola. Synonymy by Laurent (1954a:72). 
Pyxieephalus rugosus: Bocage (1870:68, 1873b:226, 1887c:211). 
Rana tuberculosa Bocage (1895a: 156, 1897b:211), Ferreira (1904:111), Schmidt (1936:130), Mertens 
(1937a: 19), Schmidt and Inger (1959:36). 
Rana {Tomopterna) tuberculosa: Monard (1938:102). 
Pyxieephalus tuberculosus: Parker (1939:142), Haacke (1970:276). 
Tomopterna signata: Cei (1997:17). 
Tomopterna rugosa: Laurent (1954a:72). 
Tomopterna tuberculosa: Loveridge (1957:345), Laurent (1964a: 133), Cei (1997:17), Poynton and Broadley 
