156 
PROCEEDINGS OF THE CALIFORNIA ACADEMY OF SCIENCES 
Series 4, Volume 65, Supplement II 
(19856:130), Poynton and Haacke (1993:13), (1996:23), Channing (2001:372), Frost (2016). 
Tomopterna (Tomopterna) tuberculosa: Fretey et al. (2011:43). 
Global conservation status (lUCN): Least Coneem. 
Global distribution: This is a widespread species occurring in savannas extending from 
Angola and northwestern Namibia across southern Democratic Republic of Congo to Tanzania, 
Zambia, and Zimbabwe. 
Ocurrences in Angola (Map 102): The 
species occurs throughout much of Angola. 
Malanje: “Pungo-Andongo” [-9.66667, 
15.58333] (Gunther 1865a:480, 1869:479; 
Boulenger 1882:30; Bocage 1895a: 156; 
Loveridge 1957:345; Schmidt and Inger 
1959:36; Poynton and Broadley 19856:130; 
Ruas 1996:23). Kwanza Sul: “Quirimbo” 
[-10.68333, 14.26667] (Parker 1936:142; Ruas 
1996:23); “Novo Redondo” [-11.20000, 
13.85000] (Ferreira 1904:111; Schmidt and 
Inger 1959:37; Ruas 1996:23). Moxico: 
“Reserve de Cameia (Cameia)” [-11.83333, 
21.00000] (Laurent 1964a:133; Ruas 1996:23). 
Bie: “Chitau” [-11.43333, 17.15000] (Schmidt 
1936:130; Ruas 1996:23); “General Machado” 
[-12.03333, 17.46667] (Mertens 1937a:19). 
Huambo: “Bimbi” [-11.81667, 15.83333] 
(Monard 1938:102; Ruas 1996:23); “Galanga” 
[-12.06667, 15.15000] (Bocage 1895a:156; Ruas 1996:23); “Mt. Moco” [-12.41667, 15.18333] 
(Parker 1936:142; Ruas 1996:23). Benguela: “Quissange” [-12.43333, 14.05000] (Bocage 
1887c:211, 1895a:156; Ruas 1996:23); “Quindumbo” [-12.46667, 14.93333] (Bocage 1895a:156; 
Ruas 1996:23); “Ebanga” [-12.73333, 14.73333] (Monard 1938:102; Ruas 1996:23); “Hanha” 
[-13.30000, 14.20000] (Bocage 1897b:211; Ruas 1996:23). Huila: “Caconda” [-13.73333, 
15.06667] (Bocage 1895a:156; Ahl “1923” 1925:45; Ruas 1996:23); “Kalukembe (Caluquembe)” 
[-13.78333, 14.68333] (Monard 1938:102; Ruas 1996:23); “13 km N of Cutenda (Cutenda 13 km 
N)” [-14.25000, 15.05000] (Poynton and Haacke 1993:13; Ruas 1996:23); “7 km N of Cutenda 
(Cutenda 7 kmN)” [-14.36667,15.08333] (Poynton and Haacke 1993:13; Ruas 1996:23); “Kuvan- 
gu” [-14.46667, 16.30000] (Monard 1938:102; Ruas 1996:23); “Huilla” [-15.05000, 13.55000] 
(Bocage 1873b:226, 1895a:156; Ahl “1923” 1925:45; Ruas 1996:23); “7 km SE of Jau (Jau 7 km)” 
[-15.21667, 13.51667] (Poynton and Haacke 1993:13); “Cascade de Ongueria (Chibia)” 
[-15.30000, 13.51667] (Laurent 1954a:72; Ruas 1996:23); “Christo Rei (Cristo Rei)” [-14.93333, 
13.51667] (Poynton and Haacke 1993:13; Ruas 1996:23). Cunene: “riusseau Mbale (Bale)” 
[-15.16667, 16.75000] (Monard 1938:102; Ruas 1996:23). Namibe: “Biballa (Bibalay’ 
[-14.76667, 13.36667] (Bocage 1895a:156; Ruas 1996:23); “Bottom of Leba Pass (Leba)” 
[-15.06667, 13.23333] (Poynton and Haacke 1993:13; Ruas 1996:23). Undetermined Locality: 
without precise locality (Bocage 1870:68). 
Taxonomic and distributional notes: The nomen Pyxicephalus rugosus was preoccupied and 
was replaced by Rana tuberculosa Boulenger, 1882. Ahl (1925 “1923”) described two new species 
of Tomopterna based on Angolan types, which Laurent (1954a) subsequently treated as junior 
synonyms of what is now T. tuberculosa. 
Map 102. Distribution of Tomopterna tuberculosa in 
Angola. 
