MARQUES ET AL.; AMPHIBIANS AND TERRESTRIAL REPTILES OF ANGOLA 
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Family Dicroglossidae Anderson, 1871 
Genus Hoplobatrachus Peters, 1863 
Hoplobatrachus occipitalis (Gunther, 1858) African Crowned Bullfrog 
Rana occipitalis Gunther 1858a:320. Syntypes: BMNH 1853.2.21.4, 1858.11.25.99-102, -112-113, 
1932.11.1.1-2 [12 specimens] (collector unknown). One of the type specimens is depicted in Gunther 
‘T858b” 1859:130, pi. 11. Type locality: “West-Afrika” (Gunther 1858a:320). 
Rana bragantina Bocage (1864:254). Holotype: MBL (collector F.A.P. Bayao), not located by Ferret 
(1967a: 19), possibly lost or destroyed by fire 18 March 1978. Type locality: “Duque de Braganga” (Boca¬ 
ge 1864:254) [= Calandula], Malanje Province, Angola. 
Rana occipitalis: Bocage (1866a:53, 1870:68, 1895a: 155), Boulenger (1882:27), Mertens (1938a:426), 
Themido (1941:2), Loveridge (1957:344), Gavetti and Andreone (1993:98), Frost (1985:508). 
Dicroglossus occipitalis: Laurent (1950a: 14, 1954a:71), Hellmich (1957a:24), Perret (1976a: 19), Cei 
(1977:16). 
Euphlyctis occipitalis: Poynton and Broadley (1985b: 125), (1996:23). 
Hoplobatrachus occipitalis: Channing (2001:295), Fretey et al. (2011:37), Channing et al. (2012:166), Frost 
(2016). 
Global conservation status (lUCN): Least Concern. 
Global distribution: The species is known from most of sub-Saharan Africa. 
Ocurrences in Angola (Map 103): The 
species known from northern and western 
Angola, though probably occurs more widely. 
Lunda Norte: “Dundo” [-7.36667, 20.83333] 
(Laurent 1954a:71); “Muita (Luembe E)” 
[-7.80000, 21.45000] (Laurent 1950a:14, 
1954a:71; Ruas 1996:23); “Muita, Luembe E, 
dans la vallee marecageuse de la Kasseke 
(afluente de la Muita)” [-7.80000, 21.45000] 
(Laurent 1950:14; Ruas 1996:23). Malanje: 
“Duque de Braganqa” [-9.10000, 15.95000] 
(Bocage 1864:254, 1866a:53, 1895a:155; Per¬ 
ret 1976a: 19; Ruas 1996:23). Kwanza Norte: 
“Ambaca” [-9.26667, 15.18333] (Bocage 
1895a:155; Ruas 1996:23); “Mucoso” 
[-9.53333, 14.65000] (Hellmich 1957a:24); 
“Dondo, sur le bord droit du Quanza” 
[-9.68333, 14.43333] (Bocage 1895a:155; 
Ruas 1996:23). Kwanza Sul: “Novo Redon¬ 
do” [-11.20000, 13.85000] (Bocage 1895a:155; 
Map 103. Distribution oi Hoplobatrachus occipitalis in 
Angola. 
Ruas 1996:23). Benguela: “Catumbella” 
[-12.43333, 13.55000] (Bocage, 1895a:155; Themido 1941:2; Gavetti and Andreone 1993:98; 
Ruas 1996:23); “Cubal” [-13.03333, 14.25000] (Mertens 1938a:426; Ruas 1996:23). Undeter¬ 
mined Locality: without precise locality (Bocage 1870:68; Laurent 1954a:71); “areas of forest and 
savanna in the north and northeast of Angola” (Cei 1977:16). 
Taxonomic and distributional notes: When publishing the description of R. bragantina, 
Bocage (1864) was unaware of Gunther’s (1858) R. occipitalis, but quickly recognized this and 
synonymized the two shortly thereafter (Bocage 1866a). 
